Herbaceous perennial, forming clonal clumps, glabrate to pilose throughout; stems erect, 15-80 cm high. Lvs narrowly obovate to oblanceolate, (3.5)-8-15-(22) cm long, imparipinnate; leaflets in 8-13 pairs, ± sessile, ovate to oblong, up to 15-(25) mm long, pilose to glabrate above and densely pilose below; teeth serrate, up to 15-(18), 1/2-3/4 to midrib. Infl. spicate, branched or not, interrupted at fruiting. Sepals 5, 1-2.5 mm long, sparsely pilose. Stamens 3-7; anthers purple. Styles 1-(2), white. Fr. 20-70, ovoid, c. 3 × 2 mm, scarcely ridged, glabrous or pilose; spines 12-55, over whole surface, ± equal and lacking a thickened conic base, 1-2-(3) mm long, red, barbed at tip.