Female flowers solitary; perianth similar to that of male flower, receptacle-tube shorter, cylindrical; staminodes 3 or 5, small or prominent; ovary spiny, the spines each terminating in a short seta; ovules many, horizontal; stigma 3-or 5-lobed.
Fruit ellipsoid to subglobose, indehiscent, fleshy, many-seeded, covered with stout conical seta-tipped fleshy spines.
Seeds elliptic in outline, subcompressed; testa smooth, hard, not bordered.
Prostrate perennial herbs or erect much-branched spiny shrubs.
Leaves simple, petiolate, or reduced to minute scales.
Tendrils solitary or paired, spiniform.
Flowers yellow, dioecious.