Petiole 2.5-5 cm, slightly pubescent when young, glabrescent; leaf blade ovate, rarely oblong-ovate or elliptic, 5-10 × 2.5-7 cm, membranous, abaxially glabrous or white pubescent when young, glabrescent, adaxially slightly pubescent when young, glabrescent, base rounded, rarely slightly truncate, margin unequally and incised-serrate, 3-lobed or undivided, apex acuminate or obtuse. Fruit 5-8; nutlets oblong, ca. 8 × 6 mm, rugose; wing oblong, slightly contracted at base, including nutlet 3-5.5 × 1-1.2 cm, wings spreading variously. Fl. Apr-May, fr. Sep.
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