Trees or shrubs; dioecious; twigs often spinescent; indumentum of simple non-glandular trichomes. Leaves alternate, petiolate; stipules inconspicuous; blades pinnately veined, eglandular, entire. Inflorescences axillary, glomerulate; flowers apetalous. Staminate flowers pedicellate; calyx-lobes 4 or 5, valvate; disc annular, thickened and fleshy; stamens 8-17, monadelphous at least at base (often appearing free in bud), the filaments slender, longer than the anthers; pollen grains 3-4-colpate, tectate, colpi operculate; pistillode minute or obsolete. Pistillate flowers long-pedicellate; calyx-lobes mostly 5 or 6, reflexed at anthesis; disc as in d; ovary usually of 3 carpels, pubescent, the ovules 1 per locule, the styles free or united at the base, deeply lacerate. Fruits capsular, dehiscing to leave a persist-ent columella; seeds smooth, ecarunculate, the endosperm fleshy, the embryo with broad thin cotyledons and short radicle.