Liana to 25 m, at base up to 15 cm ø. Leaves (sub)coriaceous, entire or up to ½ cm lobed in upper half, suborbicular to oblong-lanceolate or (ob)ovate, top (sub)obtuse to acute, up to 1½ cm acuminate, base acute-acuminate, or rounded or subcordate, (4-)5-21 by (1½-)2½-12 cm, 3-5 subplinerved or ± pinninerved; nerves 3-5(-10) pairs, arching towards the top; petiole ½-7½ cm. Glands at blade-base 2, 1-4 mm ø, entirely or largely situated on two semi-orbicular auricles 1½-5 mm ø at the apex of the petiole; blade-glands 0-4(-6), ¼-1 mm ø, submarginal in the upper half of the blade; marginal glands 0-25 at either side. Inflorescences peduncled for up to 14 cm, sometimes subsessile in short-shoots up to 25 cm, lax or condensed, in ♂ up to 150-flowered, in ♀ 2-10-flowered, tendrils 0-3, up to 4 cm long. Sterile tendrils simple or 3-fid, up to 20 cm. Bracts and bracteoles narrowly triangular, acute, ½-1 mm. Flower buds ovate, not ellipsoid. ♂ Flowers narrowly tubiform-urceolate, including the 1½-4(-8) mm long stipe 9-15 by 2-3½ mm. Hypanthium including calyx tube tubiform-urceolate, fleshy, 5½-9 mm. Calyx lobes triangular to oblong, subobtuse, 2-2½(-3) mm, reflexed. Petals oblong-lanceolate, obtuse to subacute, subentire, 2½-4 by 1-1½(-2½) mm, inserted at or near the throat of the calyx tube. Filaments 1-1½ mm, connate up to½ mm, inserted at the base of the hypanthium. Anthers 4-7 mm, ± tapering to above, subacute, ⅓-1 mm apiculate. Septa 0-⅓ mm high. Corona 0. Disk glands ½-1 mm. ♀ Flowers tubiform-campanulate, including the ¼-1 mm long stipe 5-7 by 2½-3 mm. Hypanthium including calyx tube 3-4 mm. Calyx lobes 2-4 mm. Petals 2-3 mm, inserted near the throat of the calyx tube. Staminodes c. 1 mm. Septa 0. Corona 0. Disk glands c. 1 mm. Pistil 4-6 mm; gynophore 1-2 mm. Ovary subglobose to ovoid, 1¾-2½ by 1½-2½ mm; styles ½-¾ mm, ± free; stigmas irregularly lobed, each c. 1½ mm ø. Fruit 1-2(-3?), globose to broadly ovoid, base rounded, top rounded to subacute, or fruit ± fusiform, excluding the (7-)10-30 mm long gynophore 2-6(-6½) by 1½-5 cm; pericarp coriaceous to woody,½-3 mm thick. Seeds 15-40, ± orbicular, 5-10 by 5-10 by 2½-4 mm, ± muricate and pitted; embryo 5-8 mm; cotyledons ovate to elliptic, often emarginate at one side, 4½-7½ by 4-6 mm.
Home
