Female flowers: pedicels often reflexed; sepals more or less as in the male, but longer; petals absent; disk glands 3, alternating with the carpels; ovary 3-locular, with 1 ovule per loculus; styles 3, more or less connate at the base, patent-recurved, bifid or bipartite.
Inflorescences axillary or terminal, cymose, racemose or paniculate, unisexual or rarely bisexual; male inflorescences many-flowered, female few-flowered.
Seeds ovoid-subglobose, ecarunculate; testa thinly crustaceous; albumen fleshy; cotyledons scarcely broader than the radicle.
Leaves alternate or opposite, petiolate or sessile, stipulate, sometimes stipellate, simple, entire or toothed, palminerved.
Dioecious or monoecious, glabrous annual or perennial herbs, often slender and weak.
Fruits 3-lobed, small, dehiscing septicidally into 3 bivalved cocci; endocarp thin.