Adiantum raddianum C.Presl

Delta maidenhair (en), Adiante de Raddi (fr)

Species

Pteridophytes > Polypodiales > Pteridaceae > Vittarioideae > Adiantum

Characteristics

Plants terrestrial or epilithic. Rhizome short-creeping, irregularly branched, to 50 mm long, to 2 mm in diameter, set with roots, closely spaced persistent stipe bases and scales, scales firmly chartaceous, castaneus, adnate, subulate to narrowly triangular, entire to denticulate, apex terminates in a short subulate cell, to 1 mm long, to 0.3 mm wide. Fronds to 4 mm apart, erect to arching, to 640 mm long; stipe firm, atrocastaneus, nitid, terete, to 350 mm long, to 1.2 mm in diameter, basally set with scales similar to those on rhizome, to 2 mm long, to 0.5 mm wide, glabrous apically; lamina anadromous, ovate to broadly ovate, to 3-pinnate, to 300 mm long, to 220 mm wide, with up to 12 pinna pairs; rachis and lower order axes atrocastaneus, nitid, terete, glabrous; pinnae petiolate, petiole to 25 mm long, alternate, basal pinnae largest, more widely spaced than apically, usually overlapping, to 2-pinnate, ovate, to 130 mm long, to 90 mm wide, with up to 8 pinnule pairs; pinnules petiolate, petiole to 8 mm long, alternate, usually overlapping, ovate, to 55 mm long, to 36 mm wide, with up to 5 segment pairs; ultimate segments petiolate, petioles to 2 mm long, alternate, thinly herbaceous, flabellate, narrowly to broadly cuneate, variously lobed, lobes dentate, to 10 mm long, to 10 mm wide, glabrous adaxially and abaxially. Venation evident, flabellately forked, ending in margin in sinus between teeth. Sori borne along outer margin of ultimate segment lobes, on soral flaps which veins enter, soral flaps membranous, subcircular to reniform, to 2 mm in diameter; sporangia confined to veins, sessile, capsule circular to obovate in lateral view, with 19(-21) indurated annulus cells, epistomium (4-)5-celled, hypostomium (4-)5(-6)-celled, stomium with 3 or 4 narrow cells of which walls are conspicuously thickened; Spores brown, tetrahedral-globose, trilete, rugulose, exospore (40-)44(-48) µm in equatorial diameter.
More
Rhizome shortly creeping and irregularly branched, up to 2 mm in diam. Fronds erect to arching, up to 640 mm long. Stipe terete and firm, dark brown to black and glossy, up to 350 mm long, up to 1.2 mm in diam. Lamina ovate to broadly, up to 300 x 200 mm, up to 3-pinnate, with up to 12 pinna pairs. Rachis and lower order axes dark brown and glossy. Pinnae, pinnules and segments petiolate; ultimate segments thinly herbaceous, glabrous, up to 10 x 10 mm, base mostly cuneate or narrowly cuneate, mostly longer than wide, outer margins variously lobed, outer lobe margins denticulate, venation evident, forked, ending in sinus between teeth, petioles up to 2 mm long. Sori borne along outer margin of ultimate segment lobes, up to 5 per ultimate segment. Sporangia confined to veins or near veins on membranous, near circular soral flaps up to 2 mm in diam.
Rhizome slender, short, creeping, set with castaneous, broadly lanceolate, entire rhizome-scales up to 1.5 mm long. Fronds tufted, arching, thinly herbaceous; stipe castaneous, glabrous, up to 300 mm long; lamina broadly ovate-deltate, up to 250 x 220 mm, 3-to 4-pinnate; ultimate segments obcuneate to trapeziform, up to 11 x 8 mm, with filiform petiolules, not articulated, outer margins minutely crenate-serrate, veins ending in sinuses, glabrous on both surfaces; rhachis and secondary rhachises castaneous, glabrous. Sori borne on outer margins of ultimate segments; indusial flaps subcircular to reniform, c. 1.5 mm in diameter.
Rhizomes creeping, scaly. Stipes and rachises glabrous except at very base. Laminae ovate or elliptic, 12-30 × 6-18 cm, 3-4-pinnate at base. Primary pinnae alternate, 9-12 on each side of rachis. Ultimate pinnules very variable in size and shape, generally longer than broad, narrowly obovate, obovate, broadly obovate, or rhombic, 5-17 × 3-16 mm, cuneate at base, with stalk attached ± centrally, most divided into 2 or more distinct lobes, glabrous, bright green. Indusia ± reniform with prominent sinuses, glabrous, 1-8 per pinnule on outer margin only.
Perennial with widely spreading rhizome. Fronds arching, stipe glabrous, lamina ovate-deltate, 3-or 4-pinnate, ultimate segments obcuneate, outer margins minutely cuneate-serrate, veins ending in sinuses between marginal teeth. Sori on outer margins of ultimate segments, indusial flaps reniform.
Rhizome creeping, set with rhizome scales up to 1.5 mm long. Fronds 3-or 4-pinnate. Veins of sterile pinnules ending in sinuses between marginal teeth or crenations of teeth. Ultimate segments not articulated, persistent. Sori orbicular.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention evergreen
Sexuality -
Pollination -
Spread -
Mature width (meter) 0.1 - 0.5
Mature height (meter) 0.3 - 0.55
Root system creeping-root rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light 2-6
Soil humidity 4-7
Soil texture 3-4
Soil acidity 4-8
Soil nutriment -
Hardiness (USDA) 10-12

Usage

Uses environmental use medicinal
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) 1
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Adiantum raddianum leaf picture by artaxo ines (cc-by-sa)
Adiantum raddianum leaf picture by robert eckett (cc-by-sa)
Adiantum raddianum leaf picture by Cristina Seguiti (cc-by-sa)

Distribution

Adiantum raddianum world distribution map, present in Argentina, Bolivia (Plurinational State of), Brazil, Colombia, Costa Rica, Dominica, Ecuador, Spain, Georgia, Guatemala, Indonesia, India, Jamaica, Kenya, Kiribati, Sri Lanka, Mexico, Mozambique, Mauritius, Malawi, Nicaragua, New Zealand, Peru, Portugal, Paraguay, Réunion, Saint Helena, Ascension and Tristan da Cunha, El Salvador, Sao Tome and Principe, Trinidad and Tobago, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Uruguay, United States of America, Venezuela (Bolivarian Republic of), South Africa, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:17012580-1
WFO ID wfo-0001109481
COL ID 653M9
BDTFX ID 103151
INPN ID 447953
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Adiantum mexicanum Adiantum cuneipinnulum Adiantum aemulum Adiantum colpodes Adiantum tinctum Adiantum rubellum Adiantum decorum Adiantum moorei Adiantum weigandi Adiantum decorum var. quadripinnata Adiantum werckleanum Adiantum raddianum