Shrub or coarse vine; wood soft; branches and branchlets obtusely tetragonal This content downloaded from 192.104.39.2 on Thu, 9 May 2013 15:26:14 PMAll use subject to JSTOR Terms and Conditions1973] MOLDENKE-FLORA OF PANAMA (Family 168. Verbenaceae) 117 or subterete, slight flattened at the nodes, densely or sparsely setose or long-hirsute, the hairs coarse, brownish, to 3 mm long, distinct, erect, deciduous and leaving small glandular dots on the branches; leaf-scars circular on elevated sterigmata. Leaves opposite; membranous, mostly fragile and drying dark above, lighter beneath, oblong-lanceolate to oblong or obovate, 6.5-15 cm long and 2.5-7 cm wide, entire, acute or rounded (juveniles acuminate), basally acute and some-what narrowed into the petiole in juveniles, apparently rounded or subtruncate at maturity, densely setose on both surfaces, the hairs lax, spreading, brownish, 1-4 mm long, ca. 1 mm apart, basally enlarged, denser along the larger veins and on younger leaves; petioles 3-9 mm long, densely setose. Inflorescences axillary cymes and terminal panicles, the cymes few, solitary, opposite, dense-flowered, ca. 4 cm long and 3.5 cm wide, subovate, later somewhat divaricate, the panicles pyramidal-thyrsoid to 11 cm long and wide during anthesis, dense-flowered, bracteate with greatly abbreviated sympodia, in fruit often narrow and elongate, 12-15 cm long and ca. 5 cm wide throughout, pendent, the branches ca. 14 pairs, short, few-fruiter; peduncles slender, 1.5-4 cm long, setose; pedicels slender, ca. 2 mm long, to 10 mm long in fruit, setose; bracts numerous, lanceolate, ca. 15 mm long, often concealed during anthesis by the densely congested flowers; bractlets linear, 5-8 mm long; prophylls setaceous. Flowers with the calyx campanulate or broadly obconic, lax around the corolla-tube, ca. 3.6 mm long and wide, densely setose with white divaricate hairs 2-2.5 mm long, subtruncate, 4-apiculate; corolla infundibular, the tube narrow-cylindric, ca. 5.4 mm long, ampliate above, glabrous outside, pilose in the throat, the lobes 4, oblong-lingulate, ca. 3.3 mm long and 2.6 mm wide, glabrous; stamens 4, inserted ca. 4.3 mm from the base of the corolla-tube, included in the pistillate flowers, the filaments about 0.8 mm long, densely pilose; pistil long exserted in the pistillate flowers, the style ca. 8.6 mm long, the stigma branches light, ca. 5.2 mm long, the ovary oblong, ca. 1 mm long and wide, depressed at both ends, glabrous, distinctly 4-lobed apically. Fruit yellow or orange, 15-18 mm long and wide, tetragonal, thick and fleshy, smooth, apparently 4-seeded, fruiting-calyx cupuliform, incrassate and coriaceous, ca. 4 mm long and 9 mm wide, veiny, loosely setose, basally narrow, entire.
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