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Annuals or short-lived sub-shrubby perennials. Leaves opposite or sub-opposite apically, glandular-punctate beneath. Inflorescence of loose to dense terminal corymbs. Involucre campanulate; phyllaries bi-seriate, equal; receptacle broadly conical, epaleate. Corollas tubular to funnel-shaped, tube externally glandular-stipitate, 5-lobed.. Anthers short with ovate apical appendages.. Style-arms clavate, papillate. Achenes black, oblong, 5-ribbed, with short setae on the ribs; carpopodium large, white, asymmetrical; pappus of ± 5 awned scales, or reduced, awnless and occasionally absent.
Annual or perennial herbs or shrubs. Lvs opposite, petiolate, simple, usually toothed. Infl. a corymb, usually dense. Involucral bracts in 2-(3) rows, usually subequal, mostly chartaceous. Receptacle conic; scales usually 0. Corolla tubular. Style base not swollen. Achenes ellipsoid-cylindric to obpyramidal, 5-angled, hairy; pappus usually of few free or united scales, sometimes reduced to a corona or 0.
