Tall trees. Leaves opposite and decussate, imparipinnate, 1-2(-3)-jugate with lateral leaflets subopposite, the terminal leaflets generally larger than the lateral ones, and upper lateral leaflets larger than lower ones, occasionally unifoliolate; without domatia; margin crenate, serrate to subentire. Stipules interpetiolar, one pair per node, early caducous. Inflorescence axillary, paniculate, with opposite branching pattern to outermost extremities, many-flowered, with small caducous bracts. Flowers bisexual, 4-7(-8)-merous, apetalous, with short pedicel gradually widening into discoid receptacle. Sepals valvate, coriaceous. Stamens with subulate filaments and anthers ovoid or cordate. Disc annular, ± angular following the calyx, often ± lobed with shallow radial groove inside all stamens. Ovary superior, syncarpous, with as many carpels as sepals, truncated-conical, each cell with one ovule; styles one at apex of each cell, short-conical, free, each erect or all adpressed to each other to form single conical style surface at apex of ovary. Fruit a drupe with dry-leathery exocarp and woody endocarp, the styles remaining as protruding small horns or sometimes hardly noticeable around central-apical depression. Seeds 4-8, oblong.
Home