Herbs, perennial, sometimes annual, sparingly glandular puberulent to spreading viscid-villous. Stems often reddish, 0.1-1.5 m. Leaves progressively reduced distally; distal leaves proportionately narrower than proximal; larger leaves: petiole 2-25 mm, equaling or shorter than blade; blade usually flat, sometimes undulate, 20-65 × 10-35 mm, base often oblique, obtuse, or round, margins entire or sinuate, apex acute, sometimes obtuse or round. Inflorescences: peduncle 3-25(-30) mm, involucres ovoid when mature, 4-6.5(-9) mm. Perianth deep pink to magenta, 5-15 mm. Fruits deeply convex, 2.9-4.7 × 1.5-2.8 mm; lateral ribs with 0-4 teeth, teeth usually broadly (rarely narrowly) triangular, never gland tipped, or edge of fruit wings entire or with only irregular undulations and incisions, concave side of fruit with 4-7 glands per row (glands rarely continuous or 2 rows glandless); stalks equaling or shorter than diameter of glandular head.