Alternanthera pungens Kunth

Khakiweed (en), Alternanthère (fr)

Species

Angiosperms > Caryophyllales > Amaranthaceae > Alternanthera

Characteristics

Prostrate herb, often rooting; taproot robust; stem terete, hard, on the younger parts densely clothed with appressed, minutely dentate, white hairs, 10-50 cm long. Leaves elliptic-obovate, narrowed into the petiole, with an obtuse or rounded top, on the upper surface glabrous or thinly ap-pressed-pilose, on the lower surface glabrous or on the nerves appressed-pilose, 1¼-4½ by ⅓-2 cm, in a single pair often of very unequal size; petiole 2-10 mm. Heads 1-3 in the leaf-axils, sessile, globose or oblong, white, ½-1½ cm long; bracts spine-tipped, ± 4 mm (including spine); bracteoles acuminate, very acute but not spinous, 3-4 mm long. Tepals very unequal: 2 abaxial ones much larger than the others, convex, spine-tipped, ± 5 mm long, above base right and left with hair-tuft, in the lower half with 3 upwards confluent nerves, hardening after anthesis; adaxial tepal rather flat, oblong, dentate near tip, mucronate, not spiny, at base right and left with hair-tuft, ± 3½ mm long, two inner tepals much smaller than the others, their lower halves very concave, embracing the ovary and afterwards the fruit, on the middle of their back with a patent hair-tuft. Stamens 5, all perfect; filaments (basal cup included) ½-¾ mm; anthers oblong, minute; pseudo-staminodes much shorter than filaments, broad, entire, emarginate or irregularly dentate. Style very short. Fruit falling off with perianth and bracteoles, broadly oval, much compressed, truncate or retuse, brown +-1½ mm long.
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Herbs annual. Stem diffuse, creeping, much branched, 20-30 cm tall, densely rigidly hairy. Petiole 3-10 mm, glabrous or hairy; leaf blade ovate, obovate, or elliptic-obovate, 1.5-4.5 × 0.5-1.5 cm, unequal in each pair, glabrous or annexed pilose, base acuminate, apex obtuse. Heads sessile, 1-3, axillary, white, globose or oblong, 5-10 mm. Bracts lanceolate, ca. 4 mm, spiny at apex; bracteoles lanceolate, 3-4 mm, apex acuminate, without spines. Tepals unequal, outer 2 lanceolate, ca. 5 mm, 3-veined below, rigid after anthesis, midvein stretching into spines; central segment elliptic, 3-3.5 mm, compressed; inner 2 small, enclosing ovary. Stamens 5; filaments 0.5-0.8 mm; pseudostaminodes shorter than filaments, entire or irregularly dentate. Style very short. Utricles brown, broadly ellipsoid, 1-1.5 mm. Fl. May, fr. Jul.
Herbs, perennial, 3-8 dm. Stems prostrate, villous. Leaves sessile; blade oval to obovate, usually as broad as long, 1.3-3 × 1.1-1.7 cm, apex rounded, pilose, glabrate. Inflorescences axillary, sessile; heads stramineous, globose to ovoid, 0.6-1 × 0.6-0.7 cm; bracts equaling tepals, apex attenuate. Flowers: tepals dimorphic, stramineous, lanceolate, 5-7 mm, apex acuminate, spinose tipped, sparsely villous, hairs barbed; stamens 5; anthers 3-5, globose; pseudostaminodes triangular, shorter than filaments, margins dentate. Utricles included within tepals, brown, compressed, 1.8 mm, apex truncate. Seeds lenticular, 1.3-1.5 mm.
Perennial or annual herb, ± 0.3 m high, sometimes woody at base, prostrate or erect to floating or scrambling. Leaves opposite, entire. Inflorescences in heads or short spikes, axillary, sessile or pedunculate; bracts persistent. Flowers bisexual, solitary in axils of bracts; bracteoles 2; tepals 4 or 5, glabrous or furnished with smooth or denticulate hairs. Fruit suborbicular or obovate in outline, usually compressed; capsule thin-walled or sometimes corky, indehiscent. Seeds ± lenticular.
Perennial herb. Stems procumbent. Leaves with blade elliptic to obovate, 15-45 x 3-27 mm, base cuneate to oblique. Flowers: inflorescence a dense axillary spike; perianth segments very dissimilar in form, ± 5 mm long, abaxial segments very long-aristate, awn ± 1/3 of total segment length, pungent, white; Jul.-Jun. Fruit orbicular, ± 0.2 mm long.
Prostrate, perennial herb. Leaves elliptic to obovate, base cuneate to oblique. Inflorescence a dense axillary spike. Perianth segments 5 mm long, pungent. Flowers white.
A herb. It can grow as an annual or continue growing from year to year. It lies along the ground. It grows from 1-10 cm high. It can form mats. The flowers are white.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention -
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.3 - 0.4
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c4

Environment

A tropical plant. It grows on a range of soils. It can be near creek banks and roadsides. It can grow in tropical and warm temperate places. It grows from sea level to 2,000 m above sea level. It can grow in arid places.
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Naturalized on waste land, dockyards and roadsides in lowland areas near the coast in New Guinea. In India it is a weed of pebbly soils and dry locations.
Light 4-9
Soil humidity 2-4
Soil texture 4-5
Soil acidity -
Soil nutriment -
Hardiness (USDA) 8-12

Usage

The young leaves are cooked and eaten as a vegetable.
Uses animal food environmental use food material medicinal
Edible leaves stems
Therapeutic use General tonic for rejuvenation (leaf), Abdomen (unspecified), Anodyne (unspecified), Diuretic (unspecified), Dysentery (unspecified), Fever (unspecified), Lactogogue (unspecified), Laxative (unspecified), Neuralgia (unspecified), Diaphoretic (unspecified), Abortifacient (unspecified), Calculus (unspecified), Diuretics (unspecified), Gonorrhea (unspecified), Diuretics (whole plant)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Alternanthera pungens leaf picture by susan brown (cc-by-sa)
Alternanthera pungens leaf picture by Susan Brown (cc-by-sa)
Alternanthera pungens leaf picture by susan brown (cc-by-sa)

Flower

Alternanthera pungens flower picture by Trap Hers (cc-by-sa)
Alternanthera pungens flower picture by susan brown (cc-by-sa)
Alternanthera pungens flower picture by susan brown (cc-by-sa)

Fruit

Alternanthera pungens fruit picture by susan brown (cc-by-sa)
Alternanthera pungens fruit picture by susan brown (cc-by-sa)
Alternanthera pungens fruit picture by mayoge bukapuka (cc-by-sa)

Distribution

Alternanthera pungens world distribution map, present in Angola, Argentina, Australia, Benin, Burkina Faso, Bangladesh, Bolivia (Plurinational State of), Brazil, Bhutan, Botswana, Central African Republic, China, Congo, Colombia, Cuba, Ecuador, Spain, Gabon, Guinea, Indonesia, India, Israel, Italy, Jamaica, Lao People's Democratic Republic, Sri Lanka, Mexico, Myanmar, Mozambique, Mauritius, Namibia, Pakistan, Peru, Philippines, Puerto Rico, Paraguay, Réunion, Saudi Arabia, Sudan, Somalia, South Sudan, Sao Tome and Principe, Seychelles, Chad, Togo, Thailand, Trinidad and Tobago, Uruguay, United States of America, Venezuela (Bolivarian Republic of), Viet Nam, Yemen, South Africa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:327342-2
WFO ID wfo-0000528787
COL ID C9JX
BDTFX ID 80659
INPN ID 670887
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Achyranthes leiantha Achyranthes lorentzii Alternanthera achyrantha Alternanthera procumbens Alternanthera thomensis Telanthera pungens Desmochaeta sordida Guilleminea procumbens Achyranthes mucronata Achyranthes radicans Alternanthera leiantha Bucholzia spinescens Alternanthera ciliata Alternanthera echinata Celosia echinata Illecebrum achyrantha Pityranthus crassifolius Illecebrum pungens Alternanthera repens Pupalia sordida Achyranthes nivea Achyranthes obovata Alternanthera pungens f. pauciflora Alternanthera achyrantha var. echinata Alternanthera achyrantha var. leiantha Alternanthera pungens var. leiantha Alternanthera pungens var. monocephala Alternanthera achyrantha var. pungens Alternanthera lorentzii Alternanthera repens f. monocephala Achyranthes repens Alternanthera pungens