Plants reddish brown to black. Leaves erect-spreading to squarrose, straight, panduriform, widest distally, apex symmetric; costa absent; leaf margins denticulate along leaf base; basal laminal cells rectangular to long-rectangular, marginal cells similar, walls pitted, nodose; medial laminal cells rounded-quadrate to ovate, 1-stratose entirely or sometimes 2-stratose distally, lumens rounded; laminal papillae rare, low. Sexual condition cladautoicous; perichaetial leaves differentiated, convolute-sheathing. Spores 18-28 µm.