Plants brown to black. Leaves erect, curved or secund, becoming subulate from an ovate or narrow-rectangular base, widest in proximal half of leaf, apex symmetric; costa present, percurrent and filling the leaf apices, strong, terete, usually reaching the leaf insertion; leaf margins entire or crenulate; basal laminal cells rectangular, walls thin and straight; medial laminal cells usually quadrate, 1-stratose or rarely 2-stratose juxtacostally, lumens rounded-quadrate; laminal papillae rare. Sexual condition dioicous; perichaetial leaves differentiated, convolute-sheathing. Spores (10-)13-15(-20) µm.