Plants brown to black. Leaves erect-spreading or secund, subulate-lanceolate from an ovate base, widest in proximal half of leaf, apex symmetric; costa present, percurrent and sometimes filling the leaf apices, strong but narrow, bulging abaxially, usually reaching the leaf insertion; leaf margins entire or occasionally weakly crenulate; basal laminal cells quadrate to short-rectangular, marginal cells mostly rounded-quadrate, walls weakly pitted-sinuose; medial laminal cells rounded-quadrate, 1-stratose or 2-stratose in patches or completely, lumens rounded; laminal papillae smooth to papillose. Sexual condition cladautoicous or gonioatoicous; perichaetial leaves differentiated, convolute-sheathing. Spores (40-)50-90(-110) µm.
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