Plants reddish brown to brownish green. Leaves secund to falcate-secund, mostly subulate, widest in proximal half of leaf, apex sym-metric; costa present, percurrent and filling the leaf apices, strong, terete, reaching the leaf insertion; leaf margins crenulate to strongly denticulate; basal and medial laminal cells quadrate to short-rectangular, 1-stratose, walls thin and straight, lumens smoothly rounded, quadrate to short-rectangular on the margins; laminal papillae strong, usually distant. Sexual condition dioicous; perichaetial leaves not differentiated beyond being larger than the cauline. Spores (18-)20-30(-40) µm.