Plants brown to black. Leaves erect-spreading, occasionally secund, broadly subulate from an ovate base, widest in proximal half of leaf, apex symmetric; costa present, percurrent and usually filling the leaf apices, moderately differentiated to strong, terete, reaching the leaf insertion; leaf margins entire or occasionally weakly crenulate; basal laminal cells quadrate to occasionally short-rectangular, marginal cells mostly quadrate or rounded, walls usually sinuose; medial laminal cells quadrate, 1-stratose to nearly completely 2-stratose, lumens rounded-quadrate; laminal papillae rare, low. Sexual condition clad-or gonioautoicous; perichaetial leaves differentiated, convolute-sheathing. Spores 35-60(-70) µm.
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