Plants medium-sized, in loose mats, dark to light green. Stems to 2.5(-6) cm, (0.6-)1-1.7 mm thick when dry, not or seldom branched, , primary branches erect-ascending to arcuate, julaceous when dry, ; central strand cells poorly or not differentiated; pseudoparaphyllia absent; rhizoids few. Branch leaves imbricate, appressed, or rarely slightly crisped when dry, erect to spreading, complanate, not secund when moist, broadly oblong-ligulate, sometimes slightly spatulate, , (1-)1.5-2.3(-3) mm; base broadly decurrent; margins undulate, sometimes infolded, sometimes dentate to spinulose at insertion because of very high papillae, entire at apex; apex rounded, intact; costa moderately strong, ending before apex, not obscured by laminal cells, often 2-fid at end, pellucid, abaxial costa cells papillose, papillae rounded-simple, in rows; basal laminal cells hyaline, mostly papillose, region extending to 1/3 basal leaf portion; medial and distal cells hexagonal, 7-12 µm wide, papillae many, branched. Perichaetia on terminal branches, beyond distalmost branching points, leaves with distal laminal cells papillose. Seta 0.4-1.6 cm. Capsule ovoid, urn 1.5-2.2 mm; stomata absent; annulus well developed; operculum obliquely short-rostrate, 0.3-0.6 mm; exostome teeth irregular, to 0.4 mm, not striolate proximally, papillose distally, often trabeculate; endostome very reduced, basal membrane to 2 cells high, segments irregular, fragmentary. Spores (11-)13-18(-21) µm, smooth to faintly papillose.