Anthospermum herbaceum L.F.

Species

Angiosperms > Gentianales > Rubiaceae > Anthospermum

Characteristics

Straggling, trailing or ascending perennial or perhaps sometimes annual herb 0.025–1.5(–2) m. long or tall, usually much branched at the base and usually with a ± woody tap-root; stems slender, glabrous to densely covered with short scaly hairs or with longer hairs on the young parts, reddish or purplish brown, the epidermis often peeling from the older stems, usually branched.. Leaf-blades lanceolate to ovate-lanceolate, 0.5–5.5 cm. long, 0.2–1.6 cm. wide, acute at the apex, cuneate to rounded at the base, discolorous, with small reddish spots on both surfaces, shortly scaly pubescent above and on the margins and main nerves beneath or ± glabrescent on both surfaces; petioles 1–6 mm. long; stipules with 3–5 linear fimbriae 1.5–5.5 mm. long from a base ± 2 mm. long.. Flowers several, axillary, ± sessile or pedicels under 0.5 mm. long, hermaphrodite or unisexual, monoecious, dioecious or polygamous and also to some extent dimorphic; bracts lanceolate, 1–4 mm. long, 0.8–1 mm. wide, ciliate.. Calyx-tube oblong, 1.2 mm. long, 1 mm. wide, smooth or muriculate, emarginate at the apex; lobes not developed.. Corolla white, cream, yellow or greenish; tube narrowly cylindrical or reduced, often hairy above, 2–4 mm. long in ♂ and hermaphrodite flowers, 0.5–0.6 or 3–3.5 mm. long in ♀ flowers; lobes in ♂ and hermaphrodite flowers elliptic, 2–4 mm. long, 0.5–1 mm. wide, with long hairs outside particularly near the apex; lobes in ♀ flowers either very small, 0.5 mm. long, or normal-sized, 2.6 mm. long, 0.8 mm. wide.. Stamens well exserted in ♂ and hermaphrodite flowers, the filaments projecting beyond the tube for 1–2.2 mm., absent in ♀ flowers.. Styles joined at base for 0.2–0.5 mm., the free feathery or papillate parts very variable in length, rudimentary and about 0.6–1.5 mm. long in ♂ flowers, 3.5–9 mm. long in hermaphrodite and ♀ flowers or up to 1.7 cm. long and tangled with the stems and foliage in what may be termed “super” ♀ forms, either exceeding the anthers or slightly shorter than them in hermaphrodite flowers.. Fruits straw-coloured, oblong or somewhat ovoid, 2-coccous, with a median longitudinal groove so that the general shape is like that of a pulley block, 2.1–3 mm. long, 1.5–1.8 mm. wide, smooth to densely minutely muriculate or rarely densely white hairy, each coccus with 2 ventral grooves separated by a raised ± median rib; the fruit sits on a headrest-shaped structure acute at each end, which is presumably a combination of minute pedicel and 2 small bracteoles and after the fruit has fallen this straightens out to form a thickened elliptic persistent organ ± 1.5 mm. long, 1 mm. wide, acute at both ends.. Seeds ellipsoid, 1.8–2.8 mm. long, 1.2–1.6 mm. wide, strongly compressed, ventrally excavated, soft and with a membranous brown shining finely reticulate testa, at length separating from the cocci which appear to be indehiscent.
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Perennial herb or ± subshrub, 'bisexual', 'bisexual' + 'female', 'female', 'bisexual' + 'male', 'male' + 'bisexual' + 'female', less commonly 'male' or 'male' + 'female', several-to many-stemmed, somewhat woody near base and with often ± thick, woody root; sometimes rather short-lived. Habit variable: plants scrambling, straggling or trailing and with stems to 2.5(-3) m long; sometimes ± erect, cylindrical to rounded, to c. 0.3(-0.5) m tall and with ± thick, almost rosette-1ike base; occasionally low and ± mat-or cushion-forming with shoots c. (20-)75-150(-200) mm long. Stems unbranched to much branched; branches often ± regular, arising in pairs, frequently with short branches of a higher order, often with short shoots bearing rather small leaves. Leaves decussate; blades 5-55 x (1-)2-25 mm, ± ovate, ovate-lanceolate, lanceolate to ± linear-lanceolate, cuneate to rounded at base, glabrous, ± densely papillate or shortly hairy, often distinctly discolourous; petioles 0.7-6.5 mm long; stipular sheath typically with (3-)5(-7) filiform setae ['should not be confused with small, sometimes ± linear short shoot leaves crowded in axils of long shoot leaves!'] seldom with only 1 ± triangular seta, the longest (0.3-)0.7-6.1 mm. Flowers mostly on short lateral branches, in ± sessile to somewhat elongated clusters of many to 6 (less commonly only 2) at nodes; corolla 4-merous, greenish to yellow or yellowish, sometimes reddish purplish tinged, mostly papillate or shortly hairy at least near tip. 'Bisexual', 'male': tube(1.5-)2-3.7 mm long (in 'bisexual' often longer than in 'male'), cylindrical, lobes (1.5-)2-2.7(-3.4) x 0.3-0.7 mm; anthers (0.9-)1.2-2(-2.5) mm long; 'male': small rudimentary ovary and stigmas present, the latter hidden in corolla tube; 'bisexual': gynoecium as in 'female' but stigmas only c. (2-)2.4-5(-6.4) mm long. 'Female': tube 0.3-0.7(-1.2) mm long, lobes 0.3-0.7(-1) x 0.1-0.3 mm; style ±0-0.8 mm long; stigmas 2, (2-)4-0.2 mm long; ovary 0.5-1.2 x 0.4-0.7 mm, sometimes with 4 often indistinct calyx lobes. Fruit yellowish brown or reddish brown, mostly with ±broad, conspicuous longitudinal grooves between mericarps; mericarps (1.5-)1.7-2.5(-2.8) x 0.9-1.6 mm, elliptic, oblong to ±obovate, ± densely covered with ± tuberculate structures, shortly hairy, papillate or subglabrous, occasionally with 2 obscure, ± triangular to rounded calyx lobes c. 0.2-0.3 x 0.2-0.3 mm.
Low cushion or mat-forming, perennial herb or ± erect and tufted subshrub, 0.1-0.6 m high. Leaves lanceolate, shortly hairy, base cuneate; petioles ± 0.7-6.1 mm long; stipular sheath with filiform setae, 0.3-6.1 mm long. Flowers on short lateral branches, elongated clusters at nodes. Corolla papillate near apex. Bisexual and male flowers: tube 2.0-3.7 mm long, cylindrical; anthers 1.5-2.5 mm long. Female flowers: tube 0.3-1.2 mm long. Ovary 1.2 x 0.7 mm long; style ± 0.8 mm long, stigma 2.0-10.2 mm long. Flowering time all year. Fruit yellowish brown, oblong to obovate, mericarps with conspicuous longitudinal grooves between mericarps, up to 2.8 x 1.6 mm, densely covered with tuberculate structures.
Erect, scrambling, trailing or sometimes rosette-like, perennial herb or subshrub, 0.75-3.00 m high, not dioecious. Leaves decussate; blade ovate, narrowly ovate to linear, glabrous, papillate or shortly hairy, distinctly discolorous; stipular sheath with (3-)5(-7) filiform setae, seldom with 1 triangular seta. Flowers: corolla with tube up to 3.7 mm long, lobes up to 3.4 mm long; greenish to yellow or yellowish, sometimes reddish purplish tinged; Oct.-Feb. Fruit with mericarps elliptic, oblong to obovate, covered with tuberculate structures, shortly hairy, papillate or subglabrous, mostly with broad conspicuous longitudinal grooves between mericarps.
Erect, scrambling, trailing or sometimes rosette-like perennial herb or subshrub, 0.75-3.00 m high, not dioecious. Leaves decussate, ovate, ovate-lanceolate, lanceolate to linear-lanceolate, glabrous, papillate or shortly hairy, distinctly discolorous. Stipular sheath with (3-)5(-7) filiform setae, seldom with 1 triangular seta. Mericarps elliptic, oblong to obovate, covered with tuberculate structures, shortly hairy, papillate or subglabrous, mostly with broad conspicuous longitudinal grooves between mericarps. Flowers greenish to yellow or yellowish, sometimes reddish purplish tinged.
Non-dioecious (hermaphrodite, hermaphrodite + female, female, hermaphrodite + male, male + hermaphrodite + female, less commonly male or male + female) perennial herb or ± sub shrub, several-to many-stemmed, scrambling, straggling or trailing, sometimes ± erect, occasionally low and ± mat-or cushion-forming, somewhat woody near base and often with ± thick, woody root; sometimes rather short-lived.
Hermaphrodite, male: tube (1.5)2–3.7 mm. long (in hermaphrodite often longer than in male), cylindrical, lobes (1.5)2–2.7(3.4) x 0.3–0.7 mm.; anthers (0.9)1.2–2(2.5) mm. long; male: small rudimentary ovary and stigmas present, the latter hidden in corolla tube; hermaphrodite: stigmas (2)2.4–5(6.4) mm. long, ovary 0.5–1.2 x 0.4–0.7 mm., sometimes with 4 often indistinct calyx lobes.
Fruit yellowish-brown or reddish-brown, mostly with ± broad, conspicuous longitudinal grooves between mericarps; mericarps (1.5)1.7–2.5(2.8) x 0.9–1.6 mm., elliptic, oblong to ± obovate, ± densely covered with ± tuberculate structures, shortly hairy, papillate or subglabrous, occasionally with 2 obscure, ± triangular to rounded calyx lobes 0.2–0.3 x 0.2–0.3 mm.
Leaves decussate; blades 5–55 x (1)2–25 mm., ± ovate, ovate-lanceolate, lanceolate to ± linear-lanceolate, cuneate to rounded at base, glabrous, ± densely papillate or shortly hairy, often distinctly discolourous; petioles 0.7–6.5 mm. long; stipular sheath mostly with (3)5(7) filiform setae, the longest (0.3)0.7–6.1 mm.
Stems slender, 7.5–250(300) cm. long, unbranched to much-branched; branches often ± regular, arising in pairs, frequently with short branches of a higher order, papillate, shortly hairy or sometimes glabrous, often with short shoots bearing rather small leaves.
Flowers mostly on short lateral branches, in ± sessile to somewhat elongated clusters of many to c. 6 at nodes; corolla 4-merous, greenish to yellow or yellowish, sometimes reddish-purplish tinged, mostly papillate or shortly hairy at least near tip.
Foliis fenis laevibus, caule herbaceo. Caules herbacei, diffusi, teretiusculi, rubri, laeves. Rami oppositi. Folia feno-verticillata, sessilia, lanceolata, laevia, uninervia. Flores axillares.
A fleshy herb. It can be erect or form a low mat. The leaves vary in shape and size. The flowers are in long clusters. They are greenish to yellow. The fruit are small and yellowish-brown.
Female: tube 0.3–0.7(1.2) mm. long, lobes 0.3–0.7(1) x 0.1–0.3 mm.; style 0–0.8 mm. long; stigmas (2)4–10.2 mm. long; ovary as in hermaphrodite.
Sprawling or trailing perennial to 3 m. Leaves decussate, shortly hairy, lanceolate. Flowers in elongated, axillary clusters, yellowish.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention -
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.75 - 3.0
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

It is a tropical plant. It grows between 450-2,300 m above sea level.
Light -
Soil humidity 1-3
Soil texture 7-8
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses medicinal
Edible fruits
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Distribution

Anthospermum herbaceum world distribution map, present in Burkina Faso, Eritrea, Ethiopia, Kenya, Lesotho, Mozambique, Rwanda, Saudi Arabia, eSwatini, Tanzania, United Republic of, Uganda, Yemen, South Africa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:743380-1
WFO ID wfo-0000220101
COL ID 67FBC
BDTFX ID -
INPN ID -
Wikipedia (EN)
Wikipedia (FR)

Synonyms

Anthospermum hedyotideum Anthospermum latifolium Anthospermum nodosum Anthospermum herbaceum Anthospermum lanceolatum Anthospermum mildbraedii Anthospermum muriculatum Anthospermum lanceolatum var. hedyotideum Anthospermum lanceolatum var. latifolium Anthospermum ferrugineum