Unarmed tree or shrub with simple, exstipulate pinnately lobed, irregularly incised, or entire leaves. Inflorescence a large panicle, with cymules of small flowers arranged racemosely on the branches. Pedicels articulated below the ovary. Dioecious. Male flowers: calyx 5; petals 5, imbricate; stamens 5, anthers dorsifixed. Female flowers: calyx and corolla similar to male; staminodes 5; ovary with 3 locules (2 abortive) and 1 ovule, styles 3-4, tapering from broad bases, stigmas terminal. Fruit drupe-like. Seed solitary, pendulous from a thickened funicle, 4-5-grooved; endosperm deeply ruminate.