Argythamnia P.Browne

Silverbush (en)

Genus

Angiosperms > Malpighiales > Euphorbiaceae

Characteristics

Herbs, subshrubs, or shrubs [trees], annual or perennial, monoecious or dioecious; hairs usually malpighiaceous (appressed and attached by the middle), sometimes unbranched [stellate], rarely absent; latex absent. Leaves drought deciduous or persistent, alternate, simple; stipules present, persistent or deciduous; petiole absent or present, glands usually absent (tack-shaped glands along length in A. adenophora); blade unlobed, margins entire or serrate-dentate, laminar glands absent; venation palmate (3-or 5-veined), secondary veins arcuate, not closely spaced. Inflorescences unisexual or bisexual (pistillate flowers proximal, staminate distal), axillary, racemes; glands subtending each bract 0. Pedicels present. Staminate flowers: sepals [4–]5, valvate, distinct; petals [4–]5, distinct, free or adnate to androphore, white, sometimes pale yellow-green or pale purple proximally; nectary extrastaminal, [4–]5 glands; stamens [4–](7–)10[–12] in [1–]2 whorls, connate proximally forming androphore; staminodes 0–5, at apex of androphore; pistillode absent. Pistillate flowers: sepals 5, distinct; petals usually 5, sometimes rudimentary or 0, distinct, white, sometimes pale yellow-green or pale purple proximally; nectary 5 glands; pistil 3(–4)-carpellate; styles 3, distinct or connate proximally, 2-fid, branches 6 per flower, [2 times 2-fid]. Fruits capsules, not muricate. Seeds globose to ovoid; caruncle absent [present].
More
Shrubs or herbs; monoecious (rarely dioecious); stems and foliage with in-dumentum of bifurcate (malpighiaceous) hairs, often with purplish pigment. Leaves alternate; petioles short; stipules small; blades not glandular, entire or dentate. Inflorescences short, racemiform, bisexual, with one to several basal 9 flowers; bracts small, subtending single flowers. Staminate flowers with calyx usually 5-lobed, valvate in the bud; petals 5, clawed at base, more or less adnate to base of staminal column; disc dissected, segments opposite the calyx-lobes; stamens 5-15, monadelphous, the anthers 1-2(-3) seriate, 1-3 filiform staminodia some-times present atop the column; pollen grains oblate, tectate, 3-or 4-colporate, bilaterally symmetrical; pistillode absent. Pistillate flowers with calyx 5-parted, the lobes imbricate; petals 5, imbricate, entire, sometimes reduced; disc ? dis-sected into sometimes elongated segments; ovary of 3 carpels, the ovules 1 in each locule, the styles free or basally connate, bifid, the style-branches ? dilated at the tips. Fruits capsular; seeds subglobose, foveolate to reticulate, ecarunculate, endosperm present, the embryo straight, the cotyledons broader than the radicle.
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Foliage retention deciduous
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Hardiness (USDA) 8-12

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Cultivation

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