Arytera bifoliolata S.T.Reynolds

Tree or shrub 5-10 m high, dbh c. 15 cm; bark smooth (or flaky), dark brown. Indumentum short, appressed, straight. Branchlets hairy when young; flowering twigs 1-4 mm thick. Leaves 1-jugate; petiole 0.6-5.5 cm long, hairy when young; petiolules 2-10 mm long, not to slightly 1-grooved. Leaflets opposite, ovate to elliptic (obovate), 5.3-18.6 by 1.8-7 cm, index 1.9-3.6, not to slightly falcate, coriaceous to chartaceous, (punctate), upper surface glabrous, lower surface glabrous, con-colorous with upper surface; base symmetric, acute to attenuate; apex rounded to slightly acuminate, very apex retuse to rounded (acute), not mucronu-late; venation flat above and below, concolorous with lamina to midrib reddish-brown, midrib raised below, nerves distinctly looped submarginally, veins laxly reticulate, not distinct; domatia absent. Inflorescences axillary to pseudoterminal, branch-ing basally and along rachis; rachis flattened, 2-9.5 cm long, hairy when young; first-order branches up to 3.5 cm long; cymules pleiochasial, 1-7-flowered; bracts 0.5-0.6 mm long, bracteoles 0.2-0.4 mm long; pedicels 1-5 mm long, to 7 mm in fruit, hairy. Flowers 2-3 mm in diameter. Calyx 0.9-2 mm high, deeply incised, outside hairy, inside glabrous to subpuberulous, teeth equal, not punctate, margin not membranous, apex acute. Petals 5, ovate to rhombic, 1.6-2.5 by 0.5-1.3 mm, not punctate, claw 0.2-0.5 mm long, blade gradually decurrent into claw, margin entire, pilose, apex acute to acuminate, outside pilose, inside (sub)glabrous; scales 0.5-1.1 mm long, free, basally not auriculate, (apex broadened), rather densely pilose. Disc glabrous. Stamens 7-9; filaments 2.5-4 mm long, basally pi-lose; anthers 0.5-1.1 mm long, straight, pilose, con-nective not protruding. Ovary 2-locular; style and stigma elongating to 0.8—1.5 mm in fruit, in fruit distinctly 2-lobed, upper 0.8-1.5 mm stigmatic. Fruits with 1 or 2 well-developed lobes, 0.6-1.3 by 0.5-1.7 cm, loculicidal, slightly to distinctly rugose to verrucose, subglabrous, stipe 0.5-2 mm long, broadly cunéate, lobes slightly flattened lat-erally, 6-11 by 4-7.5 mm, dorsally sharply angled to keeled; endocarp thin, chartaceous, pilose on sutures. Seeds ellipsoid to ovoid, flattened lateral-ly, c. 10 by 5 mm; arillode completely covering seed, lobed, not folded towards the base inside, thin, chartaceous, 1-layered.

Grows in lateritic soil and sandstone, in rainforest bordering mangroves.