Plants small to moderately robust, yellowish green or rarely dark green, brown proximally with age. Stems 1-5 cm. Leaves 1-8.5 × 0.7-2 mm, ovate-lanceolate to ovate, sometimes lingulate to lanceolate near stem apex, plane to somewhat concave, not undulate, only rarely with scattered abaxial teeth, apex acute, border cells and teeth with minute, verrucose or striate papillae; costa percurrent to subpercurrent, with a few abaxial teeth near apex, rarely absent; lamellae 0-3(-4), often discontinuous, 1-4 cells high; median leaf cells 35-52 µm wide, hexagonal to irregularly angled, rarely rounded, thin-walled, not or weakly collenchymatous, without trigones, smooth. Sexual condition dioicous, male plants as large as female plants or larger; perigonial bracts ovate to suborbicular, forming antheridial buds, often more than one bud per plant. Seta 1-3 per perichaetium, 0.5-3 cm. Capsule 1-3.5 × 0.5-1 mm, cylindric to ovate, usually somewhat curved, erect or often inclined; operculum 1.5-3.5 mm. Spores 10-28 µm.