Dioecious shrubs to treelets, 1-6 m tall; young branches densely tawny-tomentose, glabrous at maturity. Petiole 1-10 cm, villous, apex usually 2-glandular; leaf blade ovate or oblongelliptic to elliptic-lanceolate, 8-18 × 3-11 cm, thickly papery to thinly leathery, both surfaces densely brown-yellow pubescent when young, sparsely puberulent with age, hairs denser abaxially along veins, base cuneate, broadly cuneate, or rotund, rarely cordate, margin irregularly rough glandular serrate, apex acuminate, sometimes caudate-acuminate, or slightly falcate-inflected; lateral veins 7 or 8. Male flowers: inflorescences densely pubescent, paniculate, 7-21 × 1-2 cm; bracts small, triangular, ca. 1 mm; sepals 5 or 6, round, 1-2 mm, margins ciliate; stamens 16-20. Female flowers: inflorescences racemelike or paniculate, 3-5 cm; bracts 2-5 mm; sepals 5-7, imbricate, pubescent, 5 equal or 7 unequal, 1-2 mm, margins serrulate; ovary densely pubescent; styles 3; stigmas bifid. Persistent sepals accrescent; capsules subglobose, longitudinally 3-grooved, densely pubescent. Seeds ovoid, 3-4 mm in diam., with stripes of gray-brown spots. Fl. Aug-Sep, fr. Sep-Nov.