Basellaceae Raf.

Family

Angiosperms > Caryophyllales

Characteristics

Perennial, twining vines or trailing to climbing herbs, glabrous, often with succulent mucilaginous leaves and stems; rhizome or tubers often present. Leaves alternate, simple, entire, exstipulate. Inflorescence an axillary or terminal spike, raceme or panicle. Flowers bisexual, actinomorphic, each subtended by a bract. Bracteoles 2 (or 4 – see sepals), often adnate to perianth base. Sepals (sometimes interpreted as additional bracteoles) 2, petaloid, sometimes (not in Australia) inflated or wing-like. Perianth deeply to shallowly lobed (usually 5 segments, sometimes referred to as petaloid tepals), petals imbricate. Stamens usually 5, epitepalous; filaments free or connate in a basal ring; anthers 2-locular, versatile, dehiscing by longitudinal or apical slits or apical pores. Ovary superior, 3-carpellate, 1-locular; ovule 1, basal, campylotropous; style usually 1, often 3-lobed, sometimes 3 free styles. Fruit indehiscent, thin-walled, nut-like, surrounded and sometimes enclosed by the dry or fleshy perianth or (not in Australia) winged sepals. Seeds solitary, spherical, with a semi-circular, horseshoe-shaped or spiral embryo, endospermic or not.
More
Vines [herbs], perennial, succulent, mucilaginous, glabrous. Leaves alternate, sessile or petiolate; blade margins entire [toothed]; stipules absent. Inflorescences axillary or terminal, racemose, [spicate, or paniculate], bracteate. Flowers usually bisexual, sessile or pedicellate, subtended by 1-2 bracts; perianth hypogynous, hypanthium present; sepals 2, distinct or basally connate; petals [4-]5[-13], basally connate; nectary annular; stamens [4-]5[-9], epipetalous, basally connate; filaments erect or reflexed in bud, broadened at base; anthers [basifixed]versatile, 4-locular, dehiscence longitudinal [terminal]; pollen [6-] panto-porate; pistils 3-carpellate, unilocular; placentation basal; ovule 1; styles 3, basally connate; stigmas slender, bifid to clavate or capitate [or style 1, with stigma capitate to 3-lobed]. Fruits utricles, surrounded by dry or fleshy perianth, winged or not. Seeds usually rust colored, testa membranous; endosperm copious; embryo curved.
Subsucculent, glabrous, twining herbs with slender stems and alternate, entire, petiolate or rarely sessile, exstipulate leaves. Flowers regular, hermaphrodite or unisexual, in spikes, racemes or panicles; bracts small; bracteoles 2–4, often 2 adnate to the base of the perianth, sometimes wing-like. Perianth 5-lobed; lobes imbricate, sometimes coloured, united into a tube below or almost free, persistent. Stamens 5, opposite to the perianth-lobes, inserted at their base; filaments free, short; anthers versatile, variously dehiscing. Ovary superior, 1-locular; ovule solitary, basal, shortly stalked, campylotropous; style terminal, simple or 3-fid, or 3 free styles. Fruit indehiscent, surrounded by the persistent often fleshy perianth or winged bracteoles. Seeds solitary, almost spherical; endosperm copious or almost absent, surrounded by the spirally twisted or semi-annular embryo
Vines herbaceous or herbs twining, usually fleshy, glabrous. Leaves simple, alternate, usually petiolate, margin entire. Inflorescences of spikes, racemes, or panicles; bracts 3, caducous; bracteoles 2, persistent. Flowers bisexual, actinomorphic, small. Perianth segments 5, white or reddish, distinct or connate at base, imbricate in bud, persistent in fruit. Stamens 5, opposite to petals; filaments inserted on perianth. Ovary superior, 1-loculed; ovule 1, basal, campylotropous. Pistil united from 3 carpels. Style simple with 3 stigmas or 3 free styles. Fruit a utricle, dry or fleshy, often surrounded by persistent bracteoles and perianth. Seed globular; testa membranous; endosperm copious; embryo spirally twisted or semicircular to horseshoe-shaped.
Flowers small, actinomorphic, hermaphrodite or unisexual; bracts small; bracteoles 2–4, frequently appressed to base of perianth, sometimes winged
Flowers hermaphrodite, actinomorphic, in spikes, racemes or panicles; bracts small; bracteoles 2, often united to the base of the calyx
Stamens 5, inserted opposite and at the base of the sepals; filaments free, short; anthers with 2 parallel cells opening longitudinally
Ovary superior, 1-celled, with a solitary basal shortly stalked ovule; style terminal, often deeply divided into 3 stigmas
Seed solitary, almost spherical, with a membranous testa; endosperm copious, surrounded by the spirally twisted embryo
Ovary superior, unilocular; styles terminal, free or united or 3-fid; stigmas 3; ovule solitary, basal, short stalked
Stamens 5, inserted opposite and at base of perianth lobes; filaments free; anthers dehiscing variously
Sepals 5, often coloured, almost free or united into a 5-lobed tube, imbricate, persistent in fruit
Perianth 5-lobed, lobes united at base only or into a 5-lobed tube, imbricate, persistent in fruit
Twiners with slender steins and rather fleshy alternate entire petiolate leaves; stipules absent
Leaves alternate, entire or almost entire, usually petiolate, exstipulate
Fruit indehiscent, enveloped by persistent frequently fleshy perianth
Inflorescences of spikes, racemes or panicles, axillary or terminal
Fruit indehiscent, surrounded by the persistent often fleshy calyx
Subsucculent herbs with slender twining stems, glabrous
Seeds solitary, globose; endosperm present
Petals absent
Life form perennial
Growth form herb
Growth support -
Foliage retention -
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) -
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 8-12

Usage

Some species of Basella and Ullucus are food plants (e.g. B. alba is cultivated and used as an edible leafy vegetable); some species of Anredera are cultivated as ornamentals.
Uses ornamental
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Basellaceae unspecified picture

Distribution

Basellaceae world distribution map, present in Australia and China

Identifiers

LSID urn:lsid:ipni.org:names:77126684-1
WFO ID wfo-7000000066
COL ID 73J
BDTFX ID 100995
INPN ID 187284
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Basellaceae

Lower taxons

Basella Tournonia Ullucus Anredera