Sterile fronds pinnate, 9-21 cm long; lamina index 3-5, 7-17 by 2-4 cm, terminal segment 1.5-4 cm long, firm-herbaceous (to subcarnose?), olivaceous, with a ± terminal bulbil; rachis with a narrow wing throughout or in the upper half only; pinnae > 16 to 30, index 1-3, 1-2.4 by 0.7-0.8 cm, base usually symmetrical, cuneate, sometimes somewhat asymmetrical, its basiscopic side cuneate, the acroscopic side either broadly attenuate or with a basal acroscopic lobe, margin (bi)crenate-serrate with distinct spines in the sinuses; terminal segment usually narrowly triangular, much shorter than the remaining part of the lamina, the basal half with few lobes, tapering towards the acute or short-flagelloid apex, sometimes the whole terminal segment flagelloid; venation pattern: veins usually forming a costal areole, sometimes locally free. Fertile fronds 15-> 20 cm long; lamina index c. 10, pinnae index 4-6, 0.6-0.8 by c. 0.2 cm. Sporangia inserted all over the lower surface. Spores with a smooth cristate perispore.
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