Sterile fronds pinnate, 35-100(-120) cm long; lamina index of non-flagelloid fronds 1-3, of flagelloid fronds up to 7, 15-70(-100) by 6-30 cm, terminal segment 8-60(-70) cm long, herbaceous-pergamentaceous, green to blackish, with a (primordium of a) spherical subterminal bulbil; pinnae 8-20(-24), index (2-)3-6(-8), 4.5-22 by 1.5-4.5 cm, base +-symmetrical, angustate to broadly cuneate, margin lobed ¼-½ towards the costa, usually with a distinct tooth in each sinus, lobes usually finely serrate-crenate, sometimes entire; the two lowermost pinnae ± conform to the pinnae; terminal segment triangular; venation pattern irregular, veins forming areoles varying in shape and size, some of which with usually one, mostly excurrent included free vein, see Fig. 27i. Fertile fronds 35-85 cm long; lamina index 2-5, pinnae index 3-9,1.5-7 by 0.4-1.5(-2) cm. Sporangia inserted all over the lower surface. Spores with a smooth cristate-undulate perispore. Chromosomes n = 82, 2n = c. 120.
Usually terrestrial, sometimes low-epiphytic, in forest, 125-1650 m.
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