Sterile fronds usually pinnate, rarely simple, 10-70 cm long. Pinnate fronds 10-70 cm long; lamina index 1-3(-5), 7-40 by 3-25 cm, terminal segment 5-30 cm long, fleshy-coriaceous, olivaceous to brownish; pinnae 2-11, index 1-5, 1.5-16 by 0.8-4.5 cm, base ± symmetrical, cuneate to subcordate, margin ± entire to lobed to 1/5(-2/5) towards the costa, without or with but an incon-spicuous tooth or spine in the sinuses; terminal segment triangular, as long as to longer than the remaining part of the lamina; venation pattern: veins forming a regular network of angulate, isodiametric or elongate areoles which decrease in size towards the margin. Simple fronds 15-30 cm long, lamina index 4-7, 12-23 by 2.5-4.5 cm, base cuneate to subcordate, margin ± entire to lobed to 2/5 towards the costa, otherwise similar to the terminal segment of pinnate fronds. Fertile fronds pinnate, 20-75 cm long; lamina index 1-4, pinnae index 1-6, 1.2-9 by 0.3-1.5 cm. Sporangia inserted all over the lower surface. Spores with a smooth cristate perispore. Chromosomes n = 41.
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