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Sterile fronds pinnate, 25-90 cm long; lamina index 1-3, 20-55 by 7-30 cm, terminal segment 9-20 cm long, herbaceous to subcoriaceous, purplish or purplish-brown; pinnae 4-24, index 3-7, 5.5-17.5 by 1.5-4 cm, base ± symmetrical, narrowly to broadly cuneate, margin ± entire or finely serrate-crenate, without spines, apex acute to (long-)acuminate; terminal segment usually conform to the central pinnae, sometimes narrowly triangular and/or somewhat prolonged; venation pattern: veins forming a network of a costal areole and one to few transverse rows of smaller distal ones, part of the areoles with few to several, mostly excurrent included free veins; see Fig. 27b. Fertile fronds 30-70 cm long; pinnae index 3-8, 3-9 by 0.6-1.5 cm. Sporangia inserted mostly on and near the veins, either all over the lower surface, arranged acrostichoid, or along the margin only, arranged pteridoid. Spores smooth, undulate.
