Sterile fronds simple or pinnate, (8-)20-100 cm long. Pinnate fronds 35-100 cm long; lamina index 1-3, 25-60 by 8-40 cm, terminal segment 15-50 cm long, usually (sub)coriaceous, sometimes sub-carnose, olivaceous or brownish; pinnae 2-11, index 3-10(-20), 5-35 by (0.5-)l-7 cm, base attenuate to truncate, margin entire or irregularly sinuate especially in the lower half, without teeth or spines throughout; terminal segment ± conform to the central pinnae; venation pattern: veins anastomosing into a regular network of variously sized areoles, see Fig. 27j. Simple fronds (8-)20-75 cm long; lamina index 3-10, (7-)20-60 by (1-)3-15 cm, otherwise like the terminal segment of pinnate fronds. Fertile fronds (8-)20-90 cm long. Pinnate fronds 30-90 cm long; lamina index 1-4, pinnae index 4-10(-16), 5-17 by (0.5-)0.7-3 cm. Simple fronds (8-)20-60 cm long; lamina index 5-18, (4-)8-35 by (0.3-)0.5-4 cm. Sporangia inserted all over the lower surface. Spores with a smooth cristate perispore. Chromosomes n = 41, 2n = 122 + fragm.
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