Autoicous. Plants 1.5-4 cm long x 1-2.5 mm wide, green, yellow brown to brown when dry. Stems about 0.1-0.2 mm in diam., in cross section composed of 9-25 epidermal cells surrounding 25-70 medullary cells. Leaves normally densely imbricate, deeply concave and squarrose when moist with the lobe subvertical in position and somewhat clasping the stem, lobe suborbicular, 0.8-2 mm long, apex broadly rounded, some-times short apiculate, margins entire or ± crenulate, ventral margin broadly up curved, when spread out forming an angle of ca. 140-180° with the keel; median cells 35-50 x 25-32 µm; oil bodies homogeneous, ellipsoid-fusiform, 15-25 per cell. Lobules narrow oblong to broad obovate-triangular, 0.3-0.6 x 0.2-0.35 mm, about 1/3x lobe length, never reduced, inflated, apex oblique to truncate, not continuing into the ventral margin of the lobe, keel ± straight, free margin plane or incurved, sometimes waving, never undulate, in broad lobules making a sharp angle at the location of the second tooth (as in B. leiboldiana!), with 2(-3) widely spatiated teeth, teeth 1-3 cells long, ± inflexed, sometimes reduced. Under leaves imbricate, orbicular to broadly ovate, 0.3-0.6 x 0.4-0.8 mm, 3-5x stem width, apex rounded to truncate, margins narrowly recurved, rarely plane, bases decurrent and hardly folded, insertion line arched. Gynoecia with 1-2 innovations, bracts associated with innovations erect, obovate, shorter than perianth, apex acute or rounded, margins entire or weakly toothed, winged along the keel and with a large lobule ca. 1/3-1/2x lobe length, bracts not associated with innovation resembling vegetative leaves, without wing; bracteole margins plane or incurved, entire or slightly toothed. Perianths obovate, up to 1.2 mm long, short exserted to 1/3 or immersed, 3-keeled, dorsal surface concave or plane, the lateral keels usually winged and with1-5 teeth or lacinia up to 15 cells long, rarely smooth, the broad ventral keel rounded or with two sharp, diverging, smooth or denticulate ridges; beak about four cells long. Spores rectangular, 50-110 µm long, with about eight rudimentary rosettes; elaters 220-430 µm long.
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