Breutelia substricta (Schimp.) Magill

Plants medium-sized to large, loosely caespitose, yellowish green; terricolous or saxicolous. Stems 20-50 mm high, irregularly branched, occasionally with reddish tomentum below; in section elliptical to ± angular, central strand small, inner cortical cells in 6-10 rows, thin-walled, smaller and weakly thickened toward outside, outer cortical cells in 2-3 rows, stereids, reddish yellow, epidermal cells thin-walled, fragile, quickly broken away and stem fluted. Leaves ± crowded, rigidly appressed dry, erect-to widespreading wet, plicate only in lower leaf; ovate-to oval-lanceolate, (1.75-) 2.0-3.0 mm long, 0.5-1.0 mm wide in base; apex acuminate; margins recurved to revolute from base to upper leaf, serrulate to serrate above base. Costa short-excurrent as denticulate awn, to 0.5 mm long; superficial cells rectangular, smooth, incrassate; in section bulging dorsally, guide cells 2, large, incrassate, ventral stereid band 1(-2) cell(s) thick, exposed, dorsal stereid band strong, 3(-4) cells thick, dorsal surface cells occasionally with slightly larger lumens. Laminal cells long-rectangular, 5-10:1, evenly incrassate or strongly ana irregularly thickened with ± flexuose lumens, cells prorate at proximal end or infrequently papillae centred over lumen, basal cells long-rectangular, 10:1, weakly thickened; alar cells in large, distinct group, to 10 cells wide by 20 cells high, quadrate to short-rectangular, 1-2:1, thickened, occasionally strongly so. Dioicous. Perichaetia lateral through innovation; leaves narrowly acuminate above oval to oblong base, 2.2-2.5 mm long. Seta 7-10(-15) mm long, red-yellow; capsule ellipsoidal, striate wet or dry, 2.0-2.5 mm long, reddish yellow; exothecial cells rhomboidal, thin-walled, 2-3 rows at mouth transversely rectangular; stomata present at base of urn, subphaneropore; peristome double, light red-yellow, exostome teeth 16, narrowly triangular, 250 µm high, fragile, finely papillose, endostome rudimentary, consisting of irregular plates attached to exostome, smooth; operculum convex; spores rounded, 50 µm, surface with large wart-like plates, red-brown.