Plants medium-sized, gregarious. Stems rarely branched; central strand absent or weakly differentiated. Leaves oblong, elliptic or narrowly obovate; apex broadly acute, mucronate; margins plane to weakly incurved, crenulate distally, entire proximally; costae subpercurrent, smooth; distal laminal cells more or less rhomboidal, bulging on both exposed surfaces, but stronger adaxially; basal cells long-rectangular; marginal cells sometimes longer proximally forming a very weak border. Sexual condition dioicous. Seta very long, smooth, reddish brown to black. Capsule long-exserted, erect, furrowed; peristomate, stomata numerous; peristome double, exostome teeth long, narrow; endostome with basal membrane, segments filiform, almost as long as teeth, papillose. Operculum long conic-rostrate. Calyptra generally smooth, erose or lacerate at base. Spores small, round.