Plants large to robust, loosely caespitose, green or reddish green above, dark-green or brownish below; terricolous or corticolous. Stems 5-60 mm tall, branching by innovations, frequently tomentose; rhizoids red-brown, papillose, tubers absent. Leaves equidistant to comose, successive comal tufts frequently present, twisted dry, erect-spreading wet; elliptical, oblong-spathulate or oblong-acuminate,3-6 mm long; apex acute to acuminate; margins plane above, plane to recurved below, dentate to weakly spinose in upper 1/2 to 2/3; border absent or 1-3(-4) cells wide, generally inconspicuous. Costa short-excurrent or occasionally ending below apex, yellowish or reddish, frequently weak above; in section subrectangular, ventral surface cells present, dorsal stereid band absent, dorsal cells incrassate to substereid, hydroids present between stereid band and guide cells. Upper laminal cells rhomboidal, frequently pitted, (46-)55-75(-88) µm long, 20-25 µm wide; basal cells rectangular, pitted, occasionally reddish. Filamentous gemmae frequently produced on leaf laminae, branched, yellowish brown or reddish brown, finely papillose. Dioicous. Perichaetia terminal; leaves lanceolate to triangular, margins entire to dentate, border absent, costa aristate. Sporophytes not known.
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Plants large to robust, loosely caespitose. Stems 5-60 mm long. Leaves frequently in successive comal tufts, twisted when dry; elliptical or oblong-spathulate, 3-6 mm long; apex acute to acuminate; margins denticulate to spinose above; border 1-4 cells wide, inconspicuous; costa short-excurrent. Upper laminal cells rhomboidal, frequently pitted. Filamentous gemmae frequently produced on leaf laminae. Sporophytes not known in area.
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