Plants medium-sized to large, caespitose, green or reddish green above, brown to red-brown below; terricolous. Stems 5-15 mm tall, branching by subperichaetial innovations, tomentose below; rhizoids red-brown, coarsely papillose, tubers absent. Leaves comose, larger above, erect and frequently twisted dry, erect-spreading wet; elliptical or obovate or obovate-lanceolate, 2.4-3.7 mm long; apex acuminate; margins plane above, recurved below, obscurely denticulate above; border strong, cells in 2-4 rows, incrassate or substereids, bi-to multistratose, yellowish or occasionally reddish, in section forming multicellular knob. Costa shortly excurrent, yellowish or reddish; in section subround to round, laminal insertion ventral, ventral surface cells present, dorsal stereid band strong, stereids in 3-5 rows, dorsal surface cells incrassate, hydroids present between stereid band and guide cells. Upper laminal cells rhomboidal, frequently pitted, in conspicuous rows, (27-)42-55(-62) µm long, (11-)15-20(-25) µm wide; basal cells rectangular, pitted, pink to reddish below. Filamentous gemmae absent. Dioicous. Perichaetia terminal; leaves lanceolate to triangular, costa aristate. Seta 20-38 mm long, yellowish to red or reddish brown; capsule pendulous, straight or slightly curved, cylindrical or pyriform, contracted below mouth when dry, yellowish to red-brown, urn 2-3 mm long, neck wrinkled when dry, 1-3 mm long; exothecial cells irregular in shape, incrassate, shorter above; peristome double, exostome teeth oblong-acuminate, yellow to orange, minutely papillose, endostome segments tapered above, broadly perforated, cilia 2-3, appendiculate, basal membrane high, minutely papillose, operculum mammillate; calyptra not seen; spores round, 12-14 µm, pale yellow, essentially smooth.
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