Bryum pycnophyllum (Dixon) Mohamed

Plants medium-sized to robust, loosely caespitose to caespitose, green to yellowish green above, brownish below; terricolous or humicolous. Stems 5-45 mm tall, simple or branching by innovations, densely tomentose below; rhizoids red-brown or red, coarsely papillose, tubers generally absent. Leaves evenly spaced along stem or comose above, frequently in successive comal tufts, larger above, twisted dry, erect-spreading wet, elliptical to obovate, (2.5-)3.0-6.0 mm long; apex acute; margins plane above, recurved in lower 1/2 to 2/3, denticulate to dentate above; border 2-5 cells wide, cells sinuose, incrassate, yellowish. Costa aristate, awn to 0.7 mm long, entire to denticulate above, yellowish; in section semi-circular or subquadrate to subrectangular, ventral surface cells present, dorsal substereid band weak to strong, dorsal surface cells incrassate or substereids, hydroids present between stereid band and guide cells. Upper laminal cells short-rhomboidal, frequently narrower and incrassate towards border, pitted, (35-)40-63(-78) µm long, (18-)20-23(-28) µm wide; basal cells rectangular, pitted. Filamentous gemmae absent. Dioicous. Perichaetia terminal, leaves narrowly oblong or triangular. Seta 15-30 mm long, yellowish to reddish brown; capsule horizontal, straight to curved, yellowish or red-brown, urn 2.2-3.0 mm long, neck wrinkled dry; exothecial cells irregularly rectangular, incrassate; peristome double, exostome teeth oblong-acuminate, 43-58 µm long, apex hyaline, finely papillose, endostome segments keeled, perforated, cilia 2-4, appendiculate, basal membrane high, finely papillose; operculum conic; spores 18-22 µm, pale yellow to brown, weakly papillose.