Plants large, in thin to loose mats, light green or whitish, sometimes yellowish, dull or somewhat glossy. Stems creeping or sometimes erect, sparingly and irregularly branched, occasionally simple; hyalodermis present, sometimes indistinct, central strand sometimes present; pseudoparaphyllia absent. Stem and branch leaves similar, imbricate, , ovate to ovate-lanceolate, rarely oblong-lanceolate, not plicate; base weakly decurrent; margins plane, entire or usually serrulate to serrate at apex; apex acute to acuminate, rarely somewhat obtuse; costa double, ending just beyond base, rarely one branch reaching 1/3 leaf length; alar cells differentiated, rectangular; laminal cells papillose, . Specialized asexual reproduction absent. Sexual condition dioicous; perichaetial leaves narrowly ovate-to oblong-lanceolate, apex acute to acuminate. Seta dark red, reddish, or light brown. Capsule inclined to pendulous, cylindric, arcuate or sometimes straight, wrinkled and contracted below mouth when dry; annulus 1-or 2-seriate, deciduous, cells large; operculum conic to rostrate; peristome double; exostome teeth with fine striations between lamellae proximally, papillose distally; endostome basal membrane rather high, segments broad, keeled, cilia 2 or 3, as long or nearly as long as segments. Calyptra naked. Spores globose to ovoid, smooth or minutely papillose.