Plants medium-sized, in extensive, flat mats, green, yellow-green, or brownish, glossy. Stems creeping, , irregularly branched to subpinnate (often horizontal), ; hyalodermis absent, central strand present; pseudoparaphyllia foliose. Stem and branch leaves similar, erect or erect-spreading, ovate-lanceolate, , not plicate; base not decurrent; margins plane, entire; apex acuminate; costa double and short or occasionally ecostate; alar cells differentiated, quadrate to rectangular; laminal cells smooth; . Specialized asexual reproduction absent. Sexual condition autoicous; perichaetial leaves ovate-lanceolate, apex long-acuminate. Seta orange-red. Capsule suberect or inclined, rectangular, not or weakly curved, contracted below mouth; annulus slightly differentiated; operculum conic; peristome double; exostome teeth with external surface usually cross striolate basally; endostome basal membrane high, segments broad, keeled, cilia 1. Calyptra naked. Spores round, minutely papillose.