Perennial herbs (rarely annual) with taproot; stems prostrate to erect. Leaves simple, opposite, petiolate, ovate to lanceolate, serrate, 3-veined; base ±truncate. Capitula ±pedunculate, terminal but appearing axillary by growth of lateral shoots, solitary or in simple cymes; involucral bracts mostly 5, in 1 or 2 series, herbaceous; receptacle paleate; paleae scarious, linear-lanceolate, concavo-convex. Ray florets female and fertile; ligule 3-lobed, yellow. Disc florets bisexual and fertile, 4-merous; corolla yellow; anther sacs black; pappus of 2 (rarely 3) divergent or reflexed awns plus a ring of tiny laciniate scales. Achenes obconical, compressed, narrowly obovate or obcuneate, lacking wings; disc achenes rugose; ray achenes smooth, sometimes irregularly thickened apically. Pappus of 2 awns and usually 2 or 3 short scales.