Plants small to medium-sized, green, yellow-green, or brownish. Stems irregularly branched to pinnate; hyalodermis absent, central strand narrow; paraphyllia absent; rhizoids or rhizoid initials on stem or abaxial costa insertion, rarely forming tomentum, slightly or strongly branched, smooth; axillary hair distal cells 1-4, hyaline. Stem leaves almost erect to spreading, straight or falcate [± recurved], rounded-triangular or ovate to broadly ovate, not plicate, 1 mm; base decurrent; margins plane, or slightly recurved basally, partly entire, partly sinuate or weakly and often obtusely denticulate, limbidia absent; apex long-acuminate, ; costa single to (2/5-)1/2-4/5 leaf length, occasionally double and short, ; alar cells differentiated, transversely rectangular, rectangular, short-rectangular, or short-linear, slightly inflated, , region distinctly or indistinctly delimited, quadrate, ovate, or broadly ovate; medial laminal cells rectangular or linear; ; marginal cells 1-stratose. Sexual condition dioicous. Capsule horizontal, cylindric or short-cylindric, curved; peristome perfect; exostome margins dentate or slightly dentate distally; endostome cilia nodose or partially appendiculate. Spores 8-21 µm.
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