Plants small or medium-sized, green, yellowish green, golden brown, golden yellow, or brownish. Stems unbranched, irregularly branched, or irregularly pinnate; hyalodermis absent, central strand absent; paraphyllia absent or sometimes present; rhizoids or rhizoid initials on stem or abaxial costa insertion, rarely forming tomentum, slightly or strongly branched, smooth or slightly warty-papillose; axillary hair distal cells 1-5, hyaline. Stem leaves erect, spreading, or ± squarrose, occasionally distinctly falcate-secund, cordate, cordate-ovate, ovate, or rounded-triangular, not plicate, 1-4.6 mm; base not or hardly decurrent; margins plane, entire or slightly sinuate, limbidia absent; apex acuminate, ; costa double or single, ending below mid leaf; alar cells differentiated, rectangular, quadrate, or sometimes transversely rectangular or short-rectangular, strongly inflated, hyaline, , region distinct, quadrate, broadly ovate, ovate, or rectangular, along basal margins, ; medial laminal cells linear, rarely shortly so; marginal cells 1-stratose. Sexual condition dioicous or autoicous. Capsule horizontal, cylindric, curved; peristome perfect; exostome margins dentate or slightly dentate distally; endostome cilia usually 1-3, well developed, nodulose. Spores (10-)11-24 µm.
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