Carduus nutans L.

Nodding thistle (en), Chardon penché (fr)

Species

Angiosperms > Asterales > Asteraceae > Carduus

Characteristics

Annual or biennial. Stems branched above, (15)-45-75-(250) cm tall, with sparse cobwebby and multicellular hairs below, becoming more densely cobwebby above, ribbed; wing narrow between the deltoid to palmatifid clusters of teeth; spines 2-6 mm long. Lvs narrow-oblong, deeply pinnatifid, (5)-8-18-(40) × 2.5-10 cm, glabrescent or with scattered multicellular hairs especially on midrib and veins; basal lvs with 5-9 pairs of lobes; lobes pinnatifid to palmatifid-dentate; teeth spine-tipped; spines pale, 5-8 mm long. Capitula nodding, persistent, 3-4-(5) × 3-4-(5) cm, solitary; peduncle (0.5)-3-8-(12) cm long, (1)-2-3-(4) mm diam., leafless, not winged, with dense woolly tomentum. Involucre broadly ovoid to depressed-globose. Outer and middle involucral bracts geniculate; claw narrow-oblong, erect; limb lanceolate, usually broader than base, narrowly lanceolate-acuminate, deflexed to spreading, spine-tipped, ciliolate, the outer bracts sparsely cobwebby as well; midvein raised almost to base of limb. Inner involucral bracts linear, puberulent at base and apex; midvein not raised; apex acute, scarious, often purplish. Corolla usually red-purple, rarely white, 18-24 mm long; lobes 6-7-(8.5) mm long; swelling at throat of tube oblong, > 2 mm long. Achenes pale, ellipsoid, 3-4 × 1.5-(2) mm; pappus (12)-15-22 mm long.
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Herbs 30-100 cm tall, biennial or perennial. Stems grayish white, usually branched, sparsely cobwebby and hirsute below, densely cobwebby-felted above; wings continuous, unequally toothed; teeth triangular, margin and apex with spines. Leaves green, concolorous, hirsute along veins or sparsely cobwebby. Lower and middle cauline leaves sessile, ovate to lanceolate, (5-)10-40 × (1.5-)3-10 cm, pinnatifid or pinnatipartite; segments 5-7 pairs, obliquely triangular to triangular-ovate, margin spinulose, apex with a 4-6 mm spine. Upper cauline leaves similar but gradually smaller upward, pinnately lobed or entire. Capitula 4-6, solitary at end of stem and branches, ± nodding. Involucre ± broadly campanulate, 4-7 cm in diam. Phyllaries slightly constricted at or beyond middle then expanded into a lanceolate, spiny-tipped, and patent to recurved distal portion; outer phyllaries 14-15 × 4-5 mm; middle and inner phyllaries 1.5-2 × ca. 0.5 cm; innermost phyllaries broadly linear to linear-lanceolate, 2-2.2 × 0.2-0.3 cm, not constricted. Corolla purplish red, ca. 2.5 cm, tube ca. 1.3 cm. Achene grayish yellow with pale brown striae, ca. 3.5 mm. Pappus bristles white, unequal, to 2 cm. Fl. and fr. Jun-Sep. 2n = 16, 40.
Annuals or biennials, 40–200+ cm. Stems glabrous to tomentose; teeth of wings to 10 mm, wing spines 2–10 mm. Leaves: basal tapering to winged petioles, blades 10–40 cm, margins 1–2×-pinnately lobed; cauline sessile, shorter, margins less divided, glabrous or ± hairy. Heads borne singly or in corymbiform arrays, sometimes a few axillary, at least terminal head usually conspicuously pedunculate, often nodding, 20–40 mm. Peduncles 2–30 cm, unwinged distally or throughout, finely tomentose. Involucres hemispheric, 20–60 mm × 20–70 mm. Phyllaries lanceolate to ovate, outer and middle with appressed bases 2–4 mm wide and spreading to reflexed, appendages 2–7 mm wide, proximally glabrous or ± tomentose, distally glabrous to minutely scabridulous. spine tips 1–4 mm, inner phyllaries with unarmed, straight or twisted tips. Corollas purple, 15–28 mm, lobes 2.5–3 times longer than throat. Cypselae golden to brown, 4–5 mm; pappus bristles 13–25 mm. 2n = 16.
Herb to 1.5 m high. Stem sparsely to moderately pubescent; wings 3–10 mm wide. Leaves green, with multicellular hairs mainly along midrib and on major veins; glands absent; marginal spines 2–6 mm long, white to brownish. Capitula solitary, terminal on 10–40 cm long side-branches, nodding at maturity; involucre depressed-globose, 20–45 mm wide excluding patent bracts, persistent; median involucral bracts patent or recurved, narrowly deltate, 22–29 mm long, 2.5–3.5 mm wide at widest point, entire, sparsely arachnoid dorsally. Corolla tube 13–17 mm long; lobes 6–8 mm long. Anthers 6–7 mm long. Achenes ellipsoid, 3.3–3.8 mm long, brown, with darker longitudinal streaks. Pappus bristles 130–169, 15–20 mm long.
Biennial, 3-20 dm; lvs glabrous, or long-villous chiefly along the main veins beneath, deeply lobed, to 25 × 10 cm; heads mostly solitary and nodding at the ends of the branches, usually large, the disk (1.5-)4-8 cm wide (pressed); peduncles naked for some distance below the head; middle and outer invol bracts conspicuously broad (2-8 mm), with long, flat, spreading or reflexed, spine-pointed tip; inner bracts narrower and softer; 2n=16. Roadsides and waste places; native of Europe, now widely established in the U.S. and adj. Can., commoner westward. June-Oct. (C. thoermeri) Hybridizes with no. 2 [Carduus acanthoides L.], the hybrids partly fertile.
It is a thistle. An erect herb which keeps growing from year to year. It grows 2 m high. The leaves are grey-green. They have deep lobes and are spiny. The leaves continue down the stems as wings. The flowers are large and red or purple and drooping. They are 5-6.5 cm across. The flowers are at the ends of branches.
Life form biennial
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination entomogamy
Spread anemochory
Mature width (meter) 0.5 - 1.0
Mature height (meter) 0.7 - 1.0
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

It is a temperate plant. It grows easily on sandy sea shores. It needs an open sunny position. It is drought and frost resistant. In western China it grows in croplands and grasslands between 500-2,300 m above sea level. Tasmania Herbarium.
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Pastures, arable fields and waste places on calcareous soils.
Light 7-9
Soil humidity 3-5
Soil texture 2-5
Soil acidity 5-8
Soil nutriment -
Hardiness (USDA) 6-8

Usage

The pith of the stem is eaten after peeling and boiling. The dried flowers can be used to curdle milk.
Uses medicinal oil ornamental
Edible flowers leaves seeds stems
Therapeutic use Depurative (unspecified), Fever (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Plants are grown by seed.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -18
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Carduus nutans habit picture by Philippe Levarlet (cc-by-sa)
Carduus nutans habit picture by Hélène (cc-by-sa)
Carduus nutans habit picture by Frank Sundermeyer (cc-by-sa)

Leaf

Carduus nutans leaf picture by César Gonzalez (cc-by-sa)
Carduus nutans leaf picture by viktoria priezvisko (cc-by-sa)
Carduus nutans leaf picture by Fabrice Mélet (cc-by-sa)

Flower

Carduus nutans flower picture by val (cc-by-sa)
Carduus nutans flower picture by César Gonzalez (cc-by-sa)
Carduus nutans flower picture by зять (cc-by-sa)

Fruit

Carduus nutans fruit picture by Béatrice Parent (cc-by-sa)
Carduus nutans fruit picture by Radoslav Marinkovic (cc-by-sa)
Carduus nutans fruit picture by Yoan MARTIN (cc-by-sa)

Distribution

Carduus nutans world distribution map, present in Australia, China, France, United Kingdom of Great Britain and Northern Ireland, New Zealand, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:188890-1
WFO ID wfo-0000084791
COL ID R6ZB
BDTFX ID 13262
INPN ID 88167
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Carduus nutans Carduus nutans f. nutans Carduus nutans var. songaricus Carduus thoermeri subsp. armenus Carduus nutans var. armenus Carduus nutans var. nutans Carduus nutans var. leiophyllus

Lower taxons

Carduus nutans subsp. leiophyllus Carduus nutans subsp. macrocephalus Carduus nutans subsp. granatensis Carduus nutans subsp. platylepis Carduus nutans subsp. subacaulis Carduus nutans subsp. perspinosus Carduus nutans subsp. platypus Carduus nutans subsp. numidicus Carduus nutans subsp. micropterus Carduus nutans subsp. taygeteus Carduus nutans subsp. siculus Carduus nutans subsp. falcato-incurvus Carduus nutans subsp. trojanus Carduus nutans var. litoralis