Carex diandra Schrank

Lesser tussock-sedge (en), Laîche arrondie (fr), Laîche à deux étamines (fr), Laîche à tige arrondie (fr)

Species

Angiosperms > Poales > Cyperaceae > Carex

Characteristics

Rhizomatous; loosely tufted, pale or bright green, not tussock-forming, shoots arising singly from the rhizome. Rhizome c. 2 mm. diam., loosely covered by dark brown sheaths, roots , fibrous, chestnut-brown; shoots approximate, 2–3 mm. diam. at base including basal sheaths. Culms 15–55 cm. × c. 1 mm., wiry, trigonous, smooth below, angles scabrid above; basal sheaths dark grey-brown. Lvs < culms, 1.5–3 mm. wide, channelled, scabrid on the edges, becoming triquetrous towards the tip. Infl. 1.5–5 cm. long, a compound spike, with a few spikes distant below and ± spikes clustered above, greenish brown, rarely subtended by a lf-like bract ± = infl. Spikes c. 5 mm. long, androgynous, male fls terminal. Glumes = or slightly < utricles, ovate, brown with wide membr. margins, midrib excurrent, finely scabrid in lower glumes. Utricles 3–3.5 × 1.5–2 mm., plano-convex, occ. unequally biconvex, broadly ovoid, with short distinct nerves radiating from the base on the convex face, but smooth on the flat face, shining, dark brown; beak c. 1 mm. long, sub-pyramidal, light brown, margins scabrid, orifice bifid; stipe minute, c. 0.2 mm. long, very narrow. Stigmas 2. Nut 1.5 mm. long, biconvex or subtrigonous, obovoid, very much narrowed towards the base, dull brown.
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Stems densely clustered, 3–10 dm, aphyllopodic, rather slender above; lvs ± elongate, ± flat, mostly 1–2.5 mm wide; sheaths ventrally pale and red-dotted, prolonged 2–3 mm beyond the base of the blade; spikes androgynous, ± numerous, small, sessile, aggregated into a compact but branched infl 2–3.5(–5) cm long and seldom over 1 cm thick; bracts small, setaceous or almost like the pistillate scales; stamens 3 (in spite of the name); scales largely hyaline-scarious and stramineous or brownish, with firmer, sometimes shortly excurrent midrib; perigynia ovate or lance-ovate, 2.4–3 mm, half as wide, usually dark brown, shining, ± spreading, very firm and thick-walled, the nerveless ventral surface slightly convex and often exposed in the spike, the dorsal surface strongly convex and with a median thin, often depressed and pale strip (an extension of the dorsal suture) bordered by a pair of veins and reaching to the base; beak ± abruptly differentiated from the body of the perigynium, coarse, serrulate-margined, often pale or greenish; achene lenticular; 2n=60. Swamps, wet meadows, and Sphagnum-bogs; circumboreal, s. to N.J., Pa., Ind., Mo., Colo., and Calif.
Rhizome shortly creeping, indurate. Culms loosely tufted, 30-45[-90] cm tall, compressed trigonous, scabrous above, clothed with brown or dark brown bladeless sheaths at base. Leaves shorter than culm, blades linear, 2-3 mm wide, flat, margins scabrous, apex acuminate. Involucral bracts setaceous, shorter than inflorescence. Inflorescence spicate-paniculate, 2-3.5 cm; sometimes lowest spike ± branched; spikes androgynous, broadly ovate, 5-7 mm. Female glumes ferruginous-brown, broadly ovate, ca. 3 × 2 mm, 3-veined, margins pale hyaline, apex acuminate. Utricles ferruginous-brown, greenish above, subequaling glume, broadly ovate, plano-convex, ca. 3 × 2 mm, leathery, with 4 or 5 short veins at base abaxially, nerveless adaxially, base broadly subcordate, spongy, shortly stipitate, distal margins denticulate, apex gradually contracted into a serrulate-margined and narrowly winged beak ca. 1 mm. Nutlets tightly enveloped, obovate, plano-convex, ca. 1.4 × 1.2 mm, base broadly cuneate, shortly stipitate; style base slightly thickened; stigmas 2. Fl. and fr. Jul. 2n = 48, 50, 54, 60.
Culms (1–)2–90 cm. Leaves: sheaths adaxially whitish, conspicuously red dotted, truncate or convex at mouth, prolonged 0.4–4(–6) mm beyond base of blade; ligules 8–57 mm; foliage leaf blades 14–30(–47) cm × 1–2.5 mm. Inflorescences bisexual, straight, little interrupted, ± compound (short branch at least on most proximal node or, occasionally, unbranched) , 2–5(–6) × 0.6–1.4(–1.7) cm, proximal 0–3 branches slightly overlapping 1 above; basal branch (when present) with 2–12 spikes; proximal internode 3–7(–10) mm. Pistillate scales straw colored or brownish, 1.5–2.7 × 0.9–1.6 mm, as wide as perigynia at base, narrower distally. Perigynia ± spreading, olive to dark chestnut brown, stongly 4–6-veined and finely 2–4-veined abaxially, with narrow median strip extending full length (grooved aspect emphasized by pair of bordering veins), often membranous flap toward apex, narrowly deltoid-ovoid, unequally biconvex, (2–)2.3–2.5(–2.9) × 1–1.4 mm, shiny; beak 0.9–1.1 mm. Achenes broadly compressed-ovoid, 1.4–1.7 × 0.7–1 mm. 2n = 48, 50, 54, 60.
Shoots close together from woody, dark brown rhizomes, loosely tufted, not tussock-forming, pale or bright green, to 55 cm high. Inflorescence a spike-like greenish-brown panicle 1.5-5 cm long. Spikes many, sessile, ± densely clustered, male flowers at top. Utricles dark brown, nerved on one face; beak-margins scabrid. Stigmas 2.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality
Pollination anemogamy
Spread hydrochory
Mature width (meter) -
Mature height (meter) 0.55 - 0.9
Root system fibrous-root rhizome
Rooting depth (meter) 0.2
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Light 5-7
Soil humidity 7-9
Soil texture -
Soil acidity 3-7
Soil nutriment -
Hardiness (USDA) 4-9

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by cuttings or seedlings.
Mode cuttings seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Carex diandra habit picture by F. Manzano (cc-by-sa)

Flower

Carex diandra flower picture by F. Manzano (cc-by-sa)

Fruit

Carex diandra fruit picture by Julia Henning (cc-by-sa)

Distribution

Carex diandra world distribution map, present in Austria, Belarus, Canada, Switzerland, Chile, China, Colombia, Czech Republic, Germany, Denmark, Spain, Finland, France, Micronesia (Federated States of), Hungary, Ireland, Iran (Islamic Republic of), Iceland, Italy, Japan, Kazakhstan, Kyrgyzstan, Montenegro, Malaysia, Netherlands, Norway, New Zealand, Pakistan, Poland, Romania, Sweden, Turks and Caicos Islands, Turkmenistan, Ukraine, United States of America, and Uzbekistan

Conservation status

Carex diandra threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:299441-1
WFO ID wfo-0000346001
COL ID 69BMJ
BDTFX ID 13647
INPN ID 88468
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Carex ehrhartiana Carex bernardina Carex diandra Vignea diandra Vignea teretiuscula Carex diandra f. conguesta Carex diandra f. longibracteata Carex diandra f. major Carex diandra f. tenella Carex fulva Carex pseudoparadoxa Carex teretiuscula f. simplicior Caricina teretiuscula Carex teretiuscula Physiglochis teretiuscula Carex diandra var. ampla Carex paniculata var. diandra Carex teretiuscula var. ampla Carex teretiuscula var. crassior Carex teretiuscula var. major Carex teretiuscula var. ramosa Carex teretiuscula var. tenella Carex paniculata var. minor Carex diandra var. ramosa Carex paradoxa var. pseudoparadoxa Carex teretiuscula var. ehrhartiana Carex paradoxa subsp. pseudoparadoxa Carex teretiuscula var. nana