Carex echinata Murray

Star sedge (en), Laîche étoilée (fr), Laîche-hérisson (fr), Laîche épineuse (fr)

Species

Angiosperms > Poales > Cyperaceae > Carex

Characteristics

Rhizome short, forming dense tufts. Stems slender, erect, obtusely trigonous below, more acutely so above, smooth or slightly scaberulous just below the inflorescence, 15-90 cm by 1-2 mm. Leaves subbasal, shorter to longer than the stems, canaliculate-conduplicate, long-attenuate, scabrid on the margins above, 1-3 mm wide, the lower ones reduced to pale to castaneous, bladeless sheaths. Inflorescence ovoid to oblong, head-like or spikelike, 1.5-3.5(-4.5) cm long. Spikelets 3-8, gynaecandrous (male flowers very few), ellipsoid, obovoid or ovoid to subglobose, sessile, dense-flowered, approximate or lowest somewhat distant, 5-10 by 5-8 mm, finally squarrose by the widely spreading utricles. Bracts glumiform, the lowest with a setaceous awn, others more shortly aristate, or indistinguishable from the glumes or the lower bracts foliaceous, overtopping the inflorescence. Glumes ovate, acute, thin, wholly pale or brownish with wide whitish-hyaline margins, 3-nerved in the centre, 2.5-3.5 mm long. Utricles elliptic or ovate-lanceolate to broadly ovate, rounded to cordate at the base, plano-convex, membranous, several-nerved on both faces, glabrous, smooth, straight or slightly recurved, widely patent to reflexed when mature, spongy-thickened at the base, scarcely stipitate, (3-)4-5(-5.5) by (1-)1.5-2 mm, yellowish green to brownish, subgradually narrowed into a bidenticulate beak more or less scabrid on the margins or rarely wholly smpoth and with a dorsal split with brown overlapping margins. Nut oblong-ovoid, plano-convex, scarcely stipitate and beaked, brownish, 2-2.5 by 1-1.5 mm. Style-base slightly thickened. Stigmas 2.
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Culms 10–90(–135) cm. Leaves 3–6 per culm; sheaths tight, inner band hyaline, 1.8–2.5 cm, apex concave, glabrous; ligules 0.6–2.5(–4.5) mm, rounded to ± acute; blades plicate, 5–40 cm × 0.7–3.3(–3.8) mm, widest leaf 1–3.3(–3.8) mm wide. Inflorescences 0.7–7.8 cm; spikes (2–)3–8; lateral spikes pistillate often with few staminate flowers proximally, 3–15.5 mm, sessile, staminate portion 7-flowered, to 8.2 mm, pistillate portion 3–32-flowered, 2.5–11 mm; basal 2 spikes 1.7–42 mm apart; terminal spike gynecandrous, 5–24 mm, staminate portion 2–17-flowered, 2–16.5 × 0.9–2 mm, pistillate portion 4–26-flowered, 2.6–9 × 4.9–9.1 mm. Pistillate scales ovate, (1.4–)2.1–3.1 × 0.7–2.3 mm, apex acute to sometimes obtuse. Staminate scales ovate-lanceolate to ovate, 1.3–3.8 × 0.8–1.7 mm, base clavate, apex acute. Anthers 0.8–1.6(–2) mm. Perigynia spreading to reflexed, castaneous to dark brown, 2–14-veined abaxially, 0–12-veined adaxially over achene, lanceolate to ovate, (2.65–)2.9–4.75 × 0.8–2.1 mm, (1.7–)1.8–3.2(–3.6) times as long as wide; beak (0.85–)0.95–2 mm, (0.4–)0.45–0.86 length of body, serrulate, teeth (0–)0.15–0.5 mm. Achenes ovate-lanceolate to rhombic-ovate, 1.3–2.1 × 0.8–1.55 mm. 2n = 58.
Stems densely tufted, aphyllopodic, 1–6 dm, scabrous on the angles above; lvs 3–6 per stem, all in the basal third, to about as long as the stems, plicate, mostly 1–2.5 mm wide; spikes 3–7, sessile, small and few-fld, the terminal one with a conspicuous, slender, staminate base, some of the lateral ones often wholly pistillate; bracts small and inconspicuous; anthers 0.8–1.6 mm; perigynia 5–15, crowded, widely spreading or the lower reflexed, green or tan, planoconvex, spongy-thickened at base, lightly several-to many-nerved dorsally, few-to several-nerved or virtually nerveless ventrally, narrowly lance-triangular to move commonly lance-ovate or even ovate, 2.8–3.5(–4) mm, 1.8–3.2 times as long as wide, often serrulate-margined distally, with a prominent, slender, serrulate-margined, sharply bidentate beak 1–1.6 mm, half as long to almost as long as the body; achene lenticular; 2n=50–58. Swamps, bogs, and other wet places; circumboreal, s. to Va. (and in the mts. to N.C.), Ind., Io., Utah, and Calif. Ours is the widespread var. echinata. (C. angustior; C. cephalantha; C. josselynii; C. laricina; C. muricata, misapplied; C. stellulata)
Loosely tufted from an ascending rhizome; tufts ± flaccid. Culms (3.5)–6–35 cm. × c. 0.5 mm., subtrigonous, glab.; basal sheaths grey or light brown. Lvs us. < culms, 0.5–1.5 mm. wide, channelled to flat, margins scabrid, tapering towards the subacute tip. Infl. a compound spike 1.5–4 cm. long, green or pale brown, of 3–5 ± distant spikes, utricles widely spreading when ripe and individual spikes star-shaped; lowest spike us. subtended by a filiform scabrid bract c. ½ length of infl. Spikes androgynous, 3–7 mm. long, male fls at the base of each spike; plants occ. almost dioec. Glumes c. 1/2 length of utricle, membr., light brown, with a pale cream or green well-marked midrib and hyaline margins. Utricles 2.5–4 × c. 1.5 mm., plano-convex, us. distinctly nerved; contracted above to a narrow, dark brown beak with green, slightly or distinctly scabrid margins and slightly bifid orifice; puckered below to a rather indistinct stipe c. 2 mm. long. Stigmas 2. Nut slightly < 2 mm. long, ± biconvex, oblong-ovoid, pale brown.
Loose tufts to 35 cm high. Leaves < stems. Inflorescence 1.5-4 cm long, green or pale brown, of 3-5 ± distant sessile spikes, utricles widely spreading when ripe and individual spikes stellate. Male flowers at base of spikes. Stigmas 2.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality
Pollination anemogamy
Spread hydrochory
Mature width (meter) -
Mature height (meter) 0.35 - 0.5
Root system fibrous-root rhizome
Rooting depth (meter) 0.2
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Peat swamps, wet alpine grasslands, marshy lake shores, 1850-3600 m, locally often abundant.
Light 3-7
Soil humidity 6-8
Soil texture -
Soil acidity 4-7
Soil nutriment -
Hardiness (USDA) 4-10

Usage

Uses -
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by cuttings or seedlings.
Mode cuttings seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Carex echinata habit picture by Martin Bishop (cc-by-sa)
Carex echinata habit picture by Terje Norli (cc-by-sa)
Carex echinata habit picture by kafaktor (cc-by-sa)

Leaf

Carex echinata leaf picture by Jene T (cc-by-sa)
Carex echinata leaf picture by Jindřich Jindřich (cc-by-sa)
Carex echinata leaf picture by Terje Norli (cc-by-sa)

Flower

Carex echinata flower picture by willem (cc-by-sa)
Carex echinata flower picture by huy HO (cc-by-sa)
Carex echinata flower picture by huy HO (cc-by-sa)

Fruit

Carex echinata fruit picture by Ugoline Jacquot (cc-by-sa)
Carex echinata fruit picture by Yoan MARTIN (cc-by-sa)
Carex echinata fruit picture by Yoan MARTIN (cc-by-sa)

Distribution

Carex echinata world distribution map, present in Albania, Australia, Austria, Bulgaria, Belarus, Switzerland, Colombia, Czech Republic, Germany, Denmark, Spain, Finland, France, Micronesia (Federated States of), Greece, Guatemala, Haiti, Hungary, Indonesia, Ireland, Iran (Islamic Republic of), Iceland, Italy, Japan, Morocco, Malaysia, Nicaragua, Netherlands, Norway, New Zealand, Palau, Papua New Guinea, Poland, Korea (Democratic People's Republic of), Portugal, Romania, Russian Federation, Sweden, Turks and Caicos Islands, Ukraine, and United States of America

Conservation status

Carex echinata threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:299576-1
WFO ID wfo-0000346291
COL ID R93C
BDTFX ID 13700
INPN ID 88489
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Carex echinata f. remotiuscula Carex stellulata f. hydrophila Carex fasciculata Carex fasciculata Carex cephalantha Vignea echinata Carex echinata f. brevispicata Carex echinata f. chlorocarpa Carex serrulata Carex stellulata f. oligantha Carex echinata subsp. grypos Carex echinata var. grypos Carex echinata var. microstachys Carex echinata var. tenuior Carex hispida subsp. retusa Carex muricata var. angustata Carex muricata var. depauperata Carex muricata subsp. microcarpa Carex muricata var. monostachya Carex phyllomanica var. ormantha Carex stellulata var. australis Carex stellulata var. grypos Carex stellulata var. scirpina Carex echinata var. hydrophila Carex svenonis var. alakaiensis Carex echinata subsp. hydrophila Carex echinata subsp. gasparrinii Carex riparia subsp. fasciculata Carex echinata var. elata Carex muricata var. basilata Carex leersii var. angustata Carex stellulata f. hylogiton Carex stellulata var. subalpina Carex echinata var. echinata Carex echinata

Lower taxons

Carex echinata subsp. echinata Carex echinata subsp. phyllomanica