Carex vesicaria L.

Blister sedge (en), Laîche vésiculeuse (fr), Laîche à utricules renflés (fr)

Species

Angiosperms > Poales > Cyperaceae > Carex

Characteristics

Stems 3–10 dm, sharply trigonous, strongly scabrous above, loosely to densely clustered on a system of rather short, stout, branching rhizomes; lvs tending to be septate-nodulose, the blade elongate, flat, 3–8 mm wide; those subtending the pistillate spikes sheathless or nearly so but with elongate blade; spikes several, sessile or inconspicuously short-pedunculate, ± erect, remote, the lower pistillate, the upper staminate, or one of them androgynous; staminate spikes 2–7 cm; pistillate spikes 2–7 × 1–1.5(–2) cm at maturity; pistillate scales somewhat shorter and narrower than the perigynia, thin, acutish to long-acuminate but scarcely awned; perigynia crowded, ascending, ± in 6(–8) vertical rows, 5–8 × 2–3 mm, strongly 10–20-ribbed, lanceolate or lance-ovate in outline, bladdery-inflated below, gradually tapering to the flatter, often poorly defined beak with evident short (0.3–1.2 mm) teeth; achene yellowish, trigonous, 1.7–2.4 mm, loose in the lower part of the perigynium, continuous with the bony, eventually flexuous to strongly contorted style; 2n=74, 82. Wet soil or shallow water in bogs or swamps and the margins of ponds or streams; circumboreal, s. in Amer. to Del., Ky., Ind., Mo., and Calif. Our plants belong to the widespread var. vesicaria.
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Rhizome stoloniferous, stolons branched. Culms 30-100 cm tall, ± stiff, triquetrous, scabrous on upper part, with red-brown and bladeless sheaths at base, rudimentary sheaths usually disintegrating into reticulate fibers. Leaves slightly shorter than culm, blades 2-5 mm wide, flat, ± stiff, with transverse septate nodes between veins, sheathed. Involucral bracts leaflike, lowermost shortly sheathed, upper not sheathed. Spikes 4-6, remote; terminal 2 or 3 spikes male, approximate, linear-cylindric, 2-3.5 mm, subsessile; remaining spikes female, oblong or oblong-cylindric, 3-7 × 1-1.5 cm, densely many flowered, shortly pedunculate. Female glumes pale ferruginous or ferruginous, narrowly ovate or ovate-lanceolate, ca. 3.2 mm, membranous, 3-veined, yellow-green between veins, margins white hyaline, apex acuminate and muticous. Utricles yellowish green, obliquely patent, longer than glume, ovate or conic-ovate, inflated trigonous, 6-8 mm, subleathery, slightly nitid, glabrous, several veined, base rounded, shortly stipitate, apex attenuate into a short beak, orifice 2-toothed. Nutlets rather loosely enveloped, obovate, trigonous, 1.7-2 mm, base stipitate; style slender, usually flexuose, base not thickened; stigmas 3, rather short. Fl. and fr. May-Jul.
Plants cespitose; rhizomes short. Culms trigonous in cross section, 15–105 cm, scabrous-angled distally. Leaves: basal sheaths reddish brown to reddish purple, thickened, not spongy; ligules longer than wide; blades mid to dark green, V-to W-shaped, widest leaves 1.8–6.5 mm wide, smooth. Inflorescences 7.5–45 cm; proximal bract 10–50 cm, exceeding but not more than 2.5 times longer than inflorescence; proximal 1–3 spikes pistillate, erect or the proximal ascending, ca. 20–150-flowered, cylindric; terminal 1–3 spikes staminate, well elevated beyond summit of separate pistillate spikes. Pistillate scales lanceolate to narrowly ovate, 2.4–5.8 × 1.2–1.7 mm, shorter than perigynia, margins entire, apex acute to acuminate, awnless. Perigynia ascending, often green or straw colored, 7–12-veined, veins running into beak, ovate, (3.6–) 4–7.5(–8.2) × 1.7–3.5(–4.5) mm, 2–3.5 times as long as wide, papery, apex contracted; beak distinct, 1.1–2.6 mm, bidentulate, smooth, teeth straight, 0.3–0.9 mm. Stigmas 3. Achenes yellow to pale brown, symmetric, not indented, trigonous, smooth. 2n = 70, 74, 82, 88.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality monoecy
Pollination anemogamy
Spread barochory hydrochory
Mature width (meter) 0.5 - 1.0
Mature height (meter) 0.8 - 1.0
Root system creeping-root fibrous-root rhizome
Rooting depth (meter) 0.4
Root diameter (meter) 0.1
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Swamps, wet thickets, wet depressions in forests, marshes, sedge meadows, bogs, stream, pond, and lakeshores, often in sites inundated in spring and dry during summer; at elevations from sea level to 3,300 metres
Light 4-7
Soil humidity 7-9
Soil texture 1-6
Soil acidity 2-7
Soil nutriment -
Hardiness (USDA) 6-9

Usage

Uses animal food material
Edible leaves shoots
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by cuttings, divisions or seedlings.
Mode cuttings divisions seedlings
Germination duration (days) 14 - 28
Germination temperacture (C°) 20
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Carex vesicaria habit picture by Alain Bigou (cc-by-sa)
Carex vesicaria habit picture by Yoan MARTIN (cc-by-sa)
Carex vesicaria habit picture by Yoan MARTIN (cc-by-sa)

Leaf

Carex vesicaria leaf picture by jvanbuul (cc-by-sa)
Carex vesicaria leaf picture by sabbrent (cc-by-sa)
Carex vesicaria leaf picture by Jaroslav Bauer (cc-by-sa)

Flower

Carex vesicaria flower picture by Sander Rijfers (cc-by-sa)
Carex vesicaria flower picture by macchia1 (cc-by-sa)
Carex vesicaria flower picture by Yvon Libessart (cc-by-sa)

Fruit

Carex vesicaria fruit picture by Victor Schoenfelder (cc-by-sa)
Carex vesicaria fruit picture by alain croibien (cc-by-sa)
Carex vesicaria fruit picture by Дмитрий Дегтярев (cc-by-sa)

Distribution

Carex vesicaria world distribution map, present in Austria, Bulgaria, Belarus, Canada, Switzerland, China, Colombia, Czech Republic, Germany, Denmark, Spain, Finland, France, Micronesia (Federated States of), United Kingdom of Great Britain and Northern Ireland, Greece, Hungary, Ireland, Iran (Islamic Republic of), Italy, Kazakhstan, Kyrgyzstan, Malaysia, Netherlands, Norway, Poland, Portugal, Romania, Russian Federation, Sweden, Turks and Caicos Islands, Ukraine, and United States of America

Conservation status

Carex vesicaria threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:302885-1
WFO ID wfo-0000352391
COL ID RCHJ
BDTFX ID 14487
INPN ID 88942
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Carex furcata Carex monile Carex suilla Carex schrenkiana Carex vaseyi Trasus vesicarius Carex vesicaria f. fluitans Carex saxatilis f. dichroa Carex bongardiana Carex pendula Carex pulla f. dichroa Carex pulla f. pedunculata Carex pulla f. polystachya Carex udensis Carex vesicaria f. aurea Carex vesicaria f. brachystachys Carex vesicaria f. dichroa Carex vesicaria f. elatior Carex vesicaria f. globosa Carex vesicaria f. gracillima Carex vesicaria f. hexasticha Carex vesicaria f. monstrosa Carex vesicaria f. pendulina Carex vesicaria f. polystachya Carex vesicaria f. pungens Carex raeana Carex drymophila var. udensis Carex exsiccata var. globosa Carex exsiccata var. pungens Carex miliaris var. raeana Carex monile var. colorata Carex monile var. pacifica Carex monile var. raeana Carex pulla var. sibirica Carex salina var. ambusta Carex vesicaria var. alpigena Carex vesicaria var. brachystachys Carex vesicaria var. calcifoenum Carex vesicaria var. colorata Carex vesicaria var. dichroa Carex vesicaria var. globosa Carex vesicaria var. lanceolata Carex vesicaria var. latifolia Carex vesicaria var. pacifica Carex vesicaria var. raeana Carex vesicaria var. robusta Carex monile var. monstrosa Carex monile var. obtusisquamis Carex vesicaria var. monstrosa Carex vesicaria var. obtusisquamis Carex vesicaria subsp. alpigena Carex vesicaria subsp. virens Carex vesicaria var. virens Carex vesicaria var. fatua Carex vesicaria f. umbrosa Carex vesicaria var. subalpina Carex vesicaria var. vesicaria Carex vesicaria var. monile Carex vesicaria