Large woody climbers, without prickles. Stems tomentose, glabrescent, often bearing prominent discoid petiole scars. Leaves with lamina penniveined, sometimes with basal veins present. Inflorescences axillary, terminal or ramiflorous, subspicate or paniculate, without accrescent bracts. Male flowers: sepals c. 9–30 in vertically imbricating whorls of 3, bracteole-like, densely pilose, the outer whorls progressively smaller, the innermost whorl valvate; petals 6, equal, ± clawed; stamens 6, usually free. Female flowers: staminodes absent; carpels 6, densely pilose; style subulate, glabrous. Drupes narrowed at base into a stipe; style-scar near base; endocarp strongly curved, bony, sometimes with a dorsal ridge. Seed horseshoe-shaped; endosperm absent; cotyledons thick.