Trees; dioecious; twigs with simple hairs. Leaves alternate, petiolate, stipulate; blades pinnately veined, usually with basal glandular spots, entire. Inuforescences unisexual; c flowers in simple or compound terminal spiciform thyrses, 9 flowers in terminal few-flowered racemes; bracts eglandular. Staminate flowers subsessile, several per bract; calyx segments 3, valvate; petals absent; disc massive, pulviniform, central, ? pubescent; stamens 4-7, usually with 1 central in the disc and 1 or 2 whorls of 3 exterior to the disc; the filaments free, abruptly in-flexed near the top, the anthers introrse, dehiscing obliquely, connective pro-trusive; pollen grains subglobose (somewhat oblate), 3-colporate, tectate (tectum perforatum), triangular in polar view; pistillode absent. Pistillate flowers with stout pedicels; calyx-lobes 5 or 6, imbricate; petals absent; disc annular; ovary of 3 carpels (rarely 2 or 4). Fruits capsular, the cocci thick-walled, smooth; seeds ovoid to globose, large (over 1 cm long), endosperm present, the cotyledons broad and flat.
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