Casuarina cunninghamiana Miq.

River sheoak (en), Pin d'Australie (fr)

Species

Angiosperms > Fagales > Casuarinaceae > Casuarina

Characteristics

Tree, dioecious or rarely monoecious, 20-38 m high, with persistent, bark-covered, woody branches from which arise many, slender, flexible, deciduous, Equisetum-like, articulate branchlets, with several short, basal articles (segments) and 1-many longer, distal articles; articles with as many as 4-20 laterally rounded, longitudinal ridges. Leaves reduced to small, triangular scales, in a whorl of 4-20 at apex of each article (1 per longitudinal ridge). Flowers unisexual, wind-pollinated, much reduced, sessile, solitary in axil of a bract, grouped into unisexual inflorescences with closely spaced alternating whorls of bracts similar to scale leaves. Male inflorescences short to elongated catkin-like spikes, terminating deciduous branches. Male flowers: single stamen enclosed in bud by 1 or 2 concave perianth segments. Female inflorescences subsessile, globular heads. Female flower: perianth absent; carpels 2; ovules 2; style 2-branched and 2 filiform, well-exserted stigmas. Infructescences cone-like, globular, ± woody, formed by enlargement and thickening of accrescent bracts and bracteoles of individual flowers. Fruit a small samara, laterally compressed, apex produced into large, ± translucent wing. Seed solitary.
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Dioecious, small to large tree up to 20 m high, with branches ± erect to somewhat drooping. Young branchlets glabrous or sparsely hairy, fine (c. 0.66 mm diam.), with as many ± flat faces as lf-teeth, the sides faintly ribbed. Lf-teeth of adult branchlets usually 8, sometimes 7 or 9, or up to 11 in immature plants, narrow-triangular, yellowish toward base with a brown band at or above middle, 0.75-1 mm long; tip pale white, later withering. ♂ fls in terminal spikes 5-25-(30) mm long; whorls of bracts overlapping when fls mature. Cone globose to cylindric, rounded to flattened at apex, (5)-8-12 mm long; margins of valves obtuse or acute; seeds dull, pale whitish brown, 4-5 mm long, with pale membranous wing.
Persistent stems with 7–9 ribs, scale-leaves free to 1–2.5 mm., 0.3–0.4 mm. wide, reflexed. Deciduous branchlets 0.4–0.6 mm. diameter, with 7–9 close-set inconspicuous ribs; free part of scale-leaves 0.3–0.5(–0.6) mm. long, 0.1–0.2 mm. wide, straw-coloured with distinct dark red-brown band around united region or in apical half. ♂ inflorescence 1–1.5 mm. in diameter, whorls of bracts close-set, flowers 6–8 per whorl, bracts coloured as scale-leaves. Infructescence 8–12 mm. long, 7–10 mm. wide, ovoid; cells 6–9 per whorl, separated by small gap of ± 0.7 mm.; valve-backs with several irregularly ± longitudinal fine wrinkles; apical 2–3 mm. of valves free; samaras pale fawnish brown.. Fig. 1/1.
Tree 15-35 m high. Bark finely fissured and scaly, grey-brown. Teeth on new shoots erect. Branchlets drooping in vigorous specimens, erect in depauperate specimens; articles 4-9 mm long, 0.4-0.7 mm diam., mostly glabrous; edges of furrows often marked (when dry) by a slight ridge; phyllichnia angular to flat with a median rib; teeth 6-10, erect, 0.3-0.5 mm long, marcescent or not. Male spikes 0.4-4 cm long, 11-13 whorls per cm; anther 0.4-0.7 mm long. Cones sparsely pubescent; peduncle 2-9 mm long; cone body 7-14 mm long, 4-6 mm diam.; bracteoles broadly acute to acute. Samara 3-4 mm long.
Trees dioecious, to 25(-35) m tall, often with suckers from roots. Trunk straight, to ca. 40 cm d.b.h.; crown pyramidal; bark finely fissured and scaly, gray, adaxially pale red. Ultimate branchlets spreading to slightly pendulous, dark green, grayish green, or glaucous-green when dry, 15-38 cm × 0.5-0.7 mm; articles 4-5 mm. Leaves erect, 8(-10) per whorl, narrowly lanceolate. Male spikes 1.2-2(-4) cm. Cones ellipsoid or subglobose, 7-12 mm, truncate at both ends; apex of bracteoles acute. Samaras 3-5 mm including wing. Fl. Apr, fr. Jun-Sep. 2n = 18*.
A tall spreading tree. It grows 10-30 m high. It spreads 5-15 m across. The branches tend to droop. The leaves are reduced to rings or 8-10 tiny sharp leaf-teeth at the nodes of the stems. They are dark green. Trees are separately male and female. Male flowers are short and reddish brown spikes 3-4 cm long at the end of branches. Female flowers are red and round. They occur on mature wood. The fruit are cones which are round, grey and 1 cm across. They have sharp points on them. Plants are wind pollinated.
Deciduous branchlets 0.4–0.6 mm. in diam., with 7–9 inconspicuous ribs; scale leaves with a distinct dark red-brown band, free pan 0.3–0.5(0.6) x 0.1–0.2 mm.
Infructescences 8–12 x 7–10 mm., ovoid; valves with several irregular longitudinal fine wrinkles.
Persistent branches with 7–9 ribs; free parts of scale leaves 1–2.5 mm. long, reflexed.
Male inflorescence 1–1.5 mm. in diam.; bracts appressed, flowers 6–8 per whorl.
Subspecies 2 (1 in the flora): North America; native to ne, e Australia.
Samaras pale brown, dull.
Tree.
Life form perennial
Growth form tree
Growth support free-standing
Foliage retention evergreen
Sexuality dioecy
Pollination anemogamy
Spread -
Mature width (meter) 5.0 - 15.0
Mature height (meter) 18.0 - 27.5
Root system -
Rooting depth (meter) 0.6
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer
Photosynthetic pathway c3

Environment

Usually found by freshwater rivers and streams and the adjacent flatlands, only extending a short way up the valley slopes, mainly in alluvial sands and loams; at elevations up to 1,000 metres, but usually below 500 metres.
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Usually found by freshwater rivers and streams and the adjacent flatlands, only extending a short way up the valley slopes, mainly in alluvial sands and loams; at elevations up to 1,000 metres, but usually below 500 metres.
A tropical plant. It grows near watercourses. It grows in warm temperate places and can grow in tropical regions. In the Cairns Botanical Gardens. It can stand frost. It suits hardiness zones 9-11.
Grows along permanent streams.
Light 7-8
Soil humidity 3-6
Soil texture 3-5
Soil acidity 4-7
Soil nutriment -
Hardiness (USDA) 9-10

Usage

Uses animal food bee plant charcoal dye environmental use erosion control experimental purposes fodder food fuel fuelwood invertebrate food material ornamental paper recultivation shelter stabilisation of sandy soils tanning timber windbreak wood
Edible gums seeds
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Plants can be grown from seed. It forms suckers. It can be grown by cuttings and air-layering.
Mode cuttings seedlings
Germination duration (days) 30 - 90
Germination temperacture (C°) 18
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -7
Optimum temperature (C°) 20 - 36
Size -
Vigor -
Productivity -

Images

Habit

Casuarina cunninghamiana habit picture by Jean-Christophe Lombardo (cc-by-sa)
Casuarina cunninghamiana habit picture by Daniel Grixti-Cheng (cc-by-sa)

Leaf

Casuarina cunninghamiana leaf picture by bri (cc-by-sa)
Casuarina cunninghamiana leaf picture by Olivier Robin (cc-by-sa)
Casuarina cunninghamiana leaf picture by saad tazi (cc-by-sa)

Flower

Casuarina cunninghamiana flower picture by Zeballos Andre (cc-by-sa)
Casuarina cunninghamiana flower picture by Catherine Baldwin (cc-by-sa)
Casuarina cunninghamiana flower picture by aria salan (cc-by-sa)

Fruit

Casuarina cunninghamiana fruit picture by Zeballos Andre (cc-by-sa)
Casuarina cunninghamiana fruit picture by Pascal DUPUIS (cc-by-sa)
Casuarina cunninghamiana fruit picture by Pascal DUPUIS (cc-by-sa)

Distribution

Casuarina cunninghamiana world distribution map, present in Australia, Brazil, China, France, New Zealand, United States of America, and South Africa

Conservation status

Casuarina cunninghamiana threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:159845-1
WFO ID wfo-0000590647
COL ID RQVJ
BDTFX ID 14756
INPN ID 89308
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Casuarina cunninghamiana

Lower taxons

Casuarina cunninghamiana subsp. cunninghamiana