Male flowers: pedicels jointed; calyx closed in bud, later splitting into 2(4) unequal valvate lobes; petals 4–5, free, imbricate; disk glands 4–5(6), free, alternating with the petals, fleshy; stamens 15–35, erect in bud, borne on a convex to columnar receptacle, anthers basifixed, connective broad, thick, thecae soleiform, longitudinally dehiscent; pistillode absent.
Female flowers: pedicels stout, jointed; sepals (4)5, free, imbricate; petals 5, free, imbricate; disk hypogynous, cupuliform, fleshy; ovary 3–5-locular, with 1 ovule per loculus; styles 3–5, connate at the base, bifid or trifid.
Fruits 3-lobed to 5-lobed, dehiscing loculicidally or septicidally into 3–5 bivalved cocci; pericarp thin, crustaceous; endocarp thick, woody; columella large, persistent.
Leaves alternate, petiolate, stipulate, simple, entire, penninerved; nerves brochidodromous.
Inflorescences terminal, racemose, strobiliform when young; bracts broad, soon falling.
Stipules soon falling, leaving conspicuous subannular scars.
Dioecious shrubs or trees.
Seeds ecarunculate.
Indumentum simple.