Ceratophyllum submersum L.

Soft hornwort (en), Cératophylle submergé (fr), Cornifle submergé (fr), Cératophylle inerme (fr)

Species

Angiosperms > Ceratophyllales > Ceratophyllaceae > Ceratophyllum

Characteristics

Very similar to C. demersum in form, but usually more delicate and filamentous in appearance, differing as follows.. Leaves usually lighter green, 2–4 cm. long, branched dichotomously 3–4 times, sometimes one part of leaf branched more than another, rarely a few leaves of whorl only twice branched, usually rather fine, 0.1–0.3 mm. in diameter, but lower 1–2 segments sometimes swollen and up to 2.5 mm. across, marginal spiny teeth always minute, less than 0.1 or 0.2 mm., on lower 2 segments few or absent, few or more rarely numerous on final 2 segments.. Male flowers up to 2 mm. in diameter; mature anthers 0.6–0.8 mm. long, 0.4–0.7 mm. wide (length 1–1.5 times width).. Female perianth-segments usually longer, (1.5–)1.8–2 mm. long, 0.1–0.3 mm. wide, and relatively narrower (length (5–)7–10 times width).. Mature fruit with apical spine less than 1 up to 9 mm.; surface sometimes with few large warts or spines.
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Very similar to C. demersum in form, but usually more delicate and filamentous in appearance, differing as follows. Leaves usually lighter green, 2-4 cm long, branched dichotomously 3-4 times, sometimes one part of leaf branched more than another, rarely a few leaves of whorl only twice branched, usually rather fine, 0.1-0.3 mm in diameter, but lower 1-2 segments sometimes swollen and up to 2.5 mm across, marginal spiny teeth always minute, less than 0.1 or 0.2 mm, on lower 2 segments few or absent, few or more rarely numerous on final 2 segments. Male flowers up to 2 mm in diameter; mature anthers 0.6-0.8 mm long, 0.4-0.7 mm wide (length 1-1.5 times width). Female perianth-segments usually longer, (1.5-)1.8-2 mm long, 0.1-0.3 mm wide, and relatively narrower (length (5-)7-10 times width). Mature fruit with apical spine less than 1 up to 9 mm; surface sometimes with few large warts or spines.
Aquatic herb similar to Ceratophyllum demersum, but usually more delicate, differing as follows: Leaves usually lighter green, 2–4 cm. long, branching dichotomously 3–4 times (rarely twice in some whorls), usually rather fine, 0.1–0.3 mm. thick, to 2.5 mm. thick in lower segments, marginal teeth few, inconspicuous, often absent from lower segments, up to 0.1–0.2 mm. long.
Much like C. demersum, up to 1 m long, leaves in whorls of 6-8, adult ones 1¾-  long, bright green. Flowers not yet found in Malaysia. ♀: ripe fruit black, ± 4 mm long, only the style thickened into an apical soft spine, basal spines absent.
Male flowers to 2 mm. in diam.; anthers 0.6–0.8 x 0.4–0.7 mm., often smaller and relatively broader than in C. demersum.
Female perianth lobes (1.5) 1.8–2 x 0.1–0.3 mm., relatively narrower and usually longer than in C. demersum.
Fruit usually markedly laterally flattened with a marginal rim or wing.
Life form perennial
Growth form herb
Growth support aquatic
Foliage retention
Sexuality monoecy
Pollination hydrogamy
Spread hydrochory
Mature width (meter) -
Mature height (meter) 0.25 - 0.6
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

In places as the prec. sp., the New Gui-nean specimens were found massed in open water in Melaleuca leucadendron swamp forest, plant brown (BRASS).
Light 1-9
Soil humidity 10-12
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 5-11

Usage

Uses environmental use
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Ceratophyllum submersum unspecified picture

Distribution

Ceratophyllum submersum world distribution map, present in Anguilla, Albania, Armenia, Austria, Azerbaijan, Belgium, Bangladesh, Bulgaria, Bosnia and Herzegovina, Belarus, Botswana, Switzerland, Cameroon, Czech Republic, Germany, Denmark, Algeria, Spain, Estonia, France, United Kingdom of Great Britain and Northern Ireland, Georgia, Greece, Croatia, Hungary, Indonesia, Israel, Italy, Kazakhstan, Kyrgyzstan, Libya, Lithuania, Latvia, Morocco, North Macedonia, Myanmar, Montenegro, Malaysia, Netherlands, Oman, Papua New Guinea, Poland, Romania, Russian Federation, Saudi Arabia, Serbia, Slovakia, Slovenia, Sweden, Turkmenistan, Tunisia, Ukraine, United States of America, Uzbekistan, Viet Nam, Yemen, and South Africa

Conservation status

Ceratophyllum submersum threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:163088-1
WFO ID wfo-0000832288
COL ID SNWF
BDTFX ID 75047
INPN ID 90222
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Ceratophyllum inerme Ceratophyllum submersum Ceratophyllum laeve Ceratophyllum muticum Ceratophyllum verrucosum Ceratophyllum submersum var. granulosum Ceratophyllum demersum var. laeve Ceratophyllum submersum var. inerme Ceratophyllum submersum subsp. submersum

Lower taxons

Ceratophyllum submersum var. haynaldianum Ceratophyllum submersum var. submersum